Pison gymnopareion Pulawski, 2018

Pulawski, Wojciech J., 2018, A Revision of the Wasp Genus Pison Jurine, 1808 of Australia and New Zealand, New Guinea, and the Pacific Islands (Hymenoptera: Crabronidae), Proceedings of the California Academy of Sciences 65, pp. 1-584 : 216-217

Pison gymnopareion Pulawski , species nova

Figures 486-487.

NAME DERIVATION.– Gymnopareion is derived from two Greek words: γυμνός, naked, bare, and παρήιον, a cheek; with reference to the impunctate and glabrous lower gena (on each side of the oral cavity).

RECOGNITION.– The female of Pison gymnopareion (the male is unknown) shares with P. nudigenale a unique combination of erect setae on tergum I and the lower gena impunctate and glabrous on each side or the oral cavity, with the glabrous area bordered by a psammophore. The females of the two species are quite similar morphologically, but they differ by the shape of the clypeal lamella: in P. gymnopareion , it is evenly arcuate and markedly broader (its corners are clos- er to the adjacent orbit than to each other); in P. nudigenale , it is obtusely tridentate and relatively narrow (its corners are closer to each other than to the adjacent orbit).

DESCRIPTION.– Frons dull, finely punctate, punctures less than one diameter apart; middle supraantennal carina present, but hidden by vestiture. Occipital carina joining hypostomal carina. Gena narrow in dorsal view (as in P. nudigenale , Fig. 722 ). Mandible with well-defined abductor ridge. Labrum not emarginate. Anteromedian pronotal pit transversely elongate, about as long as midocellar diameter. Scutum not foveate along flange, without longitudinal ridges adjacent to posterior margin; scutal punctures well defined, contiguous. Tegula not enlarged. Mesopleural punctures well defined, contiguous. Postspiracular carina present, about 1.5 × as long as midocellar diameter. Metapleural sulcus costulate between dorsal and ventral metapleural pits. Propodeum with irregular longitudinal carina separating side from dorsum and posterior surface and extending from gastral socket area toward spiracle; dorsum obliquely ridged; side ridged, punctate between ridges; posterior surface irregularly transversely ridged, punctate between ridges, with several ridges radiating up from transverse carina just above gastropropodeal articulation. Posteroventral forefemoral surface with well defined punctures averaging about one diameter apart. Punctures of tergum I about one diameter apart anterior to apical depression, uniform on anterior declivity. Sterna II and III impunctate mesally, with punctures several diameters apart on each side of midline.

FIGURE 486. Pison gymnopareion Pulawski , sp. nov., Setae silvery, erect on frons, postocellar

female. Clypeus and mandibles. area, thorax, forecoxal venter, femoral venters, and tergum I; completely concealing integument on clypeus (except lamella); genal setae: see below. Apical depressions of terga with silvery, setal fasciae. Body all black. ♀.– Upper interocular distance equal to 0.60 × lower interocular distance; ocellocular distance equal to 0.9 × hindocellar diameter, distance between hindocelli equal to 0.9 × hindocellar diameter; eye height equal to 0.92 × distance between eye notches. Free margin of clypeal lamella evenly arcuate, with well-defined lateral corner (Fig. 486); corners markedly closer to orbit than to each other. Dorsal length of flagellomere I 2.8 × apical width, of flagellomere IX 1.2 × apical width. Lower gena, mandibular posterior margin, propleural and forecoxal outer margins, and forefemoral venter with psammophore (longest setae of genal, mandibular, and forefemoral psammophores about 1.1 ×, 1.0 ×, and 1.1 ×, respectively, of greatest forefemoral width); lower gena impunctate and asetose between hypostomal carina and psammophore. Mandible : trimmal carina with small incision at about two thirds of length. Length 7.0 mm; head width 2.5 mm. ♂.– Unknown. GEOGRAPHIC DISTRIBUTION (Fig. 487).– Northwestern Northern Territory, northeastern Western Australia. RECORDS.– HOLOTYPE: ♀, AUSTRALIA: Northern Territory: Victoria Highway 38.5 km SW Timber Creek at 15°42ʹ40ʺS 130°07ʹ48ʺE, 6-13 June 2001, M.E. Irwin, F.D. Parker, C. Lambkin ( ANIC). PARATYPES: AUSTRALIA: Northern Territory: same data as holotype (1 ♀, CAS). Western Australia: Drysdale River at 15°25ʹS 126°55ʹE, 3-8 Aug

FIGURE 487. Collecting localities of Pison gymnopareion 1975, I.F.B. Common and M.S. Upton (1 ♀, ANIC). Pulawski, sp. nov.

Figures

FIGURES 721-722. Pison nudigenale Pulawski, sp. nov., female. (721) Clypeus and mandibles; (722) Head in dorsal view.