Pison incurvatum Pulawski, 2018

Pulawski, Wojciech J., 2018, A Revision of the Wasp Genus Pison Jurine, 1808 of Australia and New Zealand, New Guinea, and the Pacific Islands (Hymenoptera: Crabronidae), Proceedings of the California Academy of Sciences 65, pp. 1-584 : 234-236

Pison incurvatum Pulawski , species nova

Figures 537 -540.

NAME DERIVATION.– Incurvatus (neuter: incurvatum ) is a Latin perfect passive participle of the verb incurvare, meaning curved, bent; with reference to the shape of the female clypeus.

RECOGNITION.– Pison incurvatum has two submarginal cells, the second cell short (its posterior margin is 1.2-1.5 × height), the tegula finely punctate throughout and in some wing positions completely covering the humeral plate. The male is unknown. The female can be recognized by the unique shape of the clypeus whose lamella is bent posteriorly and forms an angle with the more dorsal part ( Figs. 537, 538 ). Unlike P. aberrans , the dorsal length of flagellomere II is 2.1 × the apical widh (rather than 1.3-1.6). Additionally, and in contrast to Pison bicellulare , the scutum has short longitudinal ridges adjacent to its hindmargin. The body is either nearly all black or (in specimens from Western Australia) the legs are ferruginous

DESCRIPTION.– Frons dull, microscopically finely punctate, punctures less than one diameter apart, middle supraantennal carina at most slightly longer than midocellar diameter (mostly short- er or absent). Distance between antennal socket and orbit slightly less than socket width. Occipital carina joining hypostomal carina. Gena narrow in dorsal view. Labrum with U-shaped emargination. Anteromedian pronotal pit transversely elongate, about 4 × as long as midocellar diameter. Scutum foveate along flange, with short longitudinal ridges adjacent to posterior margin, flange slightly extending beyond scutum posterior margin; scutal punctures minute, less than one diameter apart ( Fig. 539 ). Scutellum with foveate sulcus along anterior margin. Tegula enlarged, finely punctate throughout ( Fig. 539 ), fully covering humeral plate in some wing positions. Mesopleural punctures fine, averaging about one diameter apart near center; ill-defined omaulus present in some specimens. Postspiracular carina present, about twice as long as midocellar diameter; integument depressed between postspiracular carina and episternal sulcus. Metapleuron minutely punctate, metapleural sulcus not costulate between dorsal and ventral metapleural pits. Propodeum with irregular longitudinal carina separating side from dorsum and posterior surface and extending from gastral socket area toward spiracle; dorsum regularly obliquely ridged (transversely so in posteri- or half or so in some specimens), ridges becoming markedly larger near junction with longitudinal lateral carina, median sulcus absent in some specimens; side minutely punctate, in most specimens also ridged (at least anterodorsally); posterior surface with conspicuous, transverse ridges. Forewing with two submarginal cells; second submarginal cell short (length of posterior margin 1.2-1.5 × its height). Hindcoxal dorsum with outer margin obtusely carinate. Outer surface of hindtibia with evanescent spines. Punctures of tergum I minute, averaging about one diameter apart (some punctures up to two diameters apart). Sterna minutely punctate throughout.

Setae silvery, appressed on postocellar area, gena, thorax, forecoxal venter, femoral venters, and tergum I, variously oriented on upper frons (upper frons largely asetose in some specimens); largely concealing integument on clypeus. Apical depressions of terga with silvery, setal fasciae.

Body in most specimens black, mandible ferruginous mesally, mid- and hindtibial spurs black. In specimens from Western Australia the legs are all ferruginous and the mid- and hindtibial spurs are whitish.

♀.– Upper interocular distance equal to 0.95-0.97 × lower interocular distance; ocellocular distance equal to 0.7-0.8 × hindocellar diameter, distance between hindocelli equal to 1.2-1.3 × hindocellar diameter; eye height equal to 1.04-1.10 × distance between eye notches. Apical portion of clypeal lamella bent posteriorly, forming angle with more dorsal part ( Figs. 537, 538 ). Dorsal length of flagellomere I 2.1 × apical width, of flagellomere IX 0.9 × apical width. Mandible: trimmal carina with evanescent preapical tooth, with small incision at about two thirds of length. Length 4.5-6.3 mm; head width 1.1-1.6 mm.

♂.– Unknown.

GEOGRAPHIC DISTRIBUTION (Fig. 540).– New South Wales, Northern Territory, Queensland, Tasmania, Western Australia, and also the island of New Guinea.

RECORDS.– HOLOTYPE: ♀, AUSTRALIA: Queensland: Heathlands at 11°45ʹS 142°35ʹE 18 Sept –21 Oct 1992, P. Zborowski and T. Weir ( ANIC).

PARATYPES: AUSTRALIA: New South Wales: Dunns Swamp , 13 Nov 1982, N.W. Rodd (1 ♀, AMS) . Northern Territory: Keep River National Park at 15°44ʹ17ʺS 129°06ʹ55ʺE, 7-8 June 2001, M.E. Irwin, F.D. Parker, and Ch. Lambkin (1 ♀, 1 ♂, CAS) , at 15°45ʹ30ʺS 129°06ʹ28ʺE, 3-17 June 2001, E.T. Weir, K. Pullen, and P. Bouchard (3 ♀, ANIC) , 6-9 June 2001, Ch. Lambkin, M.E. Irwin, and F.D. Parker (3 ♀, CAS) , and at 15°55ʹ17ʺS 129°03ʹ31ʺE, 2-10 June 2001, M.E. Irwin, F.D. Parker, and Ch. Lambkin (1 ♀, CAS) . Queensland: Cockatoo Creek crossing 17 km NW Heathlands at 11°39ʹS 142°27ʹE, 22 Mar – 25 Apr 1992, T. McLeod (1 ♀, CAS) ; Gunshot Creek 13 km NW Heathlands Homestead at 11°43ʹS 142°28ʹE, 20 Mar 1992, G. Daniels and M.A Schneider (1 ♀, QMB) ; Heathlands at 11°45ʹS 142°35ʹE, June – 25 July 1992, P. Zborowski and E.S. Nielsen (1 ♀, ANIC) ; 12 km SSE Heathlands at 11°51ʹS 142°38ʹE, 22 Mar – 25 Apr 1992, T. McLeod (1 ♀, CAS) ; 14 km ENE Heathlands at 11°41ˈS 142°42ˈE, 19 Mar 1994, P. Zborowski (1 ♀, ANIC) ; 3 km ENE Mount Tozer at 12°44ʹS 143°14ʹE, 28 June – 4 July 1986, J. C. Cardale (1 ♀, ANIC) ; 3 km NE Mount Webb at 15°03ˈS 145°09ˈE, 1-30 Oct 1980, J.C. Cardale (1 ♀, ANIC) ; Shiptons Flats at 15°47ʹS 145°14ʹE, 17-19 Oct 1980, J.C. Cardale (1 ♀, ANIC) . Tasmania: 9 km SE Miena , 10 Jan 1992, D.W. Webb (1 ♀, UCD) . Western Australia: Hamersley Station at 22°29.10ʹS 117°41.28ʹE, 30 Sept – 5 Oct 2004, CVA [= Conservation Volunteers Australia] (3 ♀, AMS) ; Nanutarra-Wittenoom road at 22°26.8ʹS 117°49.56ʹE, CVA [= Conservation Volunteers Australia], 30 Sept – 5 Oct 2004 (1 ♀, AMS) , 18-23 Nov 2004 (1 ♀, CAS) , and 28 Oct – 2 Nov 2005 (1 ♀, AMS) .

INDONESIA: Western Papua (= Indonesian New Guinea): 11 km S Bupul at 7º39ʹS 140º53ʹE (2 ♀, RMNH); Erambu 80 km NE Merauke (1 ♀, BISH) .

Figures

FIGURES 537-539. Pison incurvatum Pulawski, sp. nov., female. (537) Clypeus and mandible in frontal view; (538) Clypeus obliquely from below (arrow shows bent posteriorluy apical part of clypeal lamella); (539) Tegula and adjacent scutum. FIGURE 540. Collecting localities of Pison incurvatum Pulawski, sp. nov.