Pison emarginatum Pulawski, 2018

Pulawski, Wojciech J., 2018, A Revision of the Wasp Genus Pison Jurine, 1808 of Australia and New Zealand, New Guinea, and the Pacific Islands (Hymenoptera: Crabronidae), Proceedings of the California Academy of Sciences 65, pp. 1-584 : 169-171

publication ID	https://doi.org/ 10.5281/zenodo.13159946
persistent identifier	https://treatment.plazi.org/id/E62387EA-FF1C-FF19-410D-FCDFFDAFF865
treatment provided by	Felipe
scientific name	Pison emarginatum Pulawski
status	sp. nov.

Treatment

Pison emarginatum Pulawski , species nova

Figures 357 View FIGUES -364.

NAME DERIVATION.– Emarginatum, Latin neuter for emarginate, with reference to the markedly emarginate male sternum VIII.

RECOGNITION.– Pison emarginatum has a black gaster, three submarginal cells, the second recurrent vein interstitial with the second intersubmarginal vein or nearly so, and the setae appressed on tergum I. Also, the ferruginous tibiae are correlated with the absence of the longitudinal carina separating the propodeal side from the dorsum and posterior surface, one of four such species. Unlike the other three species with this character combination ( P. aurifex , P. elongatum , and P. pilifrons ), the frons of P. emarginatum is swollen just above the antennal socket ( Fig. 359 View FIGUES ) rather than not swollen, the female has an angulate (rather than rounded) clypeal lamella, and male sternum VIII is deeply emarginate apically ( Fig. 361 View FIGURES ) rather than truncate or rounded, narrowly emarginate mesally in pilifrons . The minutely punctate metapleuron (markedly finer than the mesopleuron and the adjacent part of the propodeum) is a subsidiary recognition feature. Unlike P. pilifrons , the punctures of the upper frons are well defined (rather than microscopically small, practically unrecognizable), the mesopleural interspaces are shiny, only slightly microsculptured (rather than dull, conspicuously microsculptured), and the setae of the scutum are appressed (rather than erect).

DESCRIPTION.– Frons swollen just above antennal socket ( Fig. 359 View FIGUES ), dull, finely punctate, punctures less than one diameter apart. Labrum not emarginate. Pronotal collar roundly angulate laterally. Anteromedian pronotal pit transversely elongate, about as long as midocellar diameter. Scutum not foveate along flange, without longitudinal ridges adjacent to posterior margin; scutal punctures minute, less than one diameter apart. Tegula slightly enlarged. Mesopleural punctures larger than those on scutum, about one diameter apart near center ( Fig. 360 View FIGUES ), up to about three diameters apart in specimen from Victoria. Postspiracular carina present, as long as or shorter than midocellar diameter. Metapleuron microscopically punctate (markedly finer than mesopleuron and adjacent part of propodeum); metapleural sulcus not costulate between dorsal and ventral metapleural pits. Propodeum without longitudinal carina separating side from dorsum and posterior surface in male from Black Mountain, Australian Capital Territory, the carina is replaced by series of short, transverse carinae); dorsum obliquely ridged, punctate between ridges; side punctate, minutely ridged behind spiracle; posterior surface ridged, punctate between ridges. Posteroventral forefemoral surface finely punctate, punctures 2-3 diameters apart. Hindcoxal dorsum with outer margin not carinate in anterior half. Punctures of tergum I about one diameter apart on horizontal portion, compressed just adjacent to apical depression. Sternum II punctate throughout, punctures well defined, about 2-3 diameters apart (about 1-2 diameters apart in specimen from Canberra).

Setae golden (silvery in specimens from Canberra, Victoria, and Whiskers, New South Wales), appressed on scutum and tergum I; setae of lower gena subappressed, slightly curved, about as long as midocellar diameter; largely concealing integument on clypeus in female, completely so in male. Apical depressions of terga with golden setal fasciae (but fasciae silvery in specimen from Canberra and that from Victoria).

Head, thorax, propodeum, and gaster black; antenna ferruginous, black apically (all black in specimen from Canberra and that from Victoria). Femora black basally, ferruginous apically (all black in specimen from Canberra), tibiae, and tarsi ferruginous.

♀.– Upper interocular distance equal to 0.76 × lower interocular distance; ocellocular distance equal to 1.0-1.4 × hindocellar diameter, distance between hindocelli equal to 0.8-1.1 × hindocellar diameter; eye height equal to 1.00-1.04 × distance between eye notches. Free margin of clypeal lamella angulate ( Fig. 357 View FIGUES ). Dorsal length of flagellomere I 1.8-2.0 × apical width, of flagellomere IX 1.3 × apical width. Length 9.9-14.6 mm; head width 2.5-3.3 mm.

♂.– Upper interocular distance equal to 0.80-0.86 × lower interocular distance; ocellocular distance equal to 1.2-1.7 × hindocellar diameter, distance between hindocelli equal to 1.2 × hindocellar diameter; eye height equal to 1.00-1.08 × distance between eye notches. Free margin of clypeal lamella acutely angulate ( Fig. 358 View FIGUES ). Dorsal length of flagellomere I 1.9 × apical width, of flagellomere X 1.3 × apical width. Sternum VIII deeply emarginate apically ( Fig. 361 View FIGURES ). Genitalia: Figs. 362, 363 View FIGURES . Length 8.5-11.1 mm; head width 2.4-2.8 mm.

GEOGRAPHIC DISTRIBUTION (Fig. 364).– Eastern New South Wales, eastern Queensland, Victoria.

RECORDS.– HOLOTYPE: ♂, AUSTRALIA: New South Wales: Clarence in Blue Mountains , 28 Jan 1987, N.W. Rodd ( AMS).

PARATYPES: AUSTRALIA: Australian Capital Territory: Black Mountain , 11-31 Dec 1979, D.H. Colless (1 ♂, ANIC) ; Farrer, southern suburb of Canberra at 35°22ʹS 149°05ʹE, 1 Jan 1988, D.C.F. Rentz (1 ♀, ANIC) . New South Wales: Lake George Cullerin , 15 Feb 1988, M.E. Irwin (4 ♂, UCD) ; 0.5 km SE Lansdowne near Taree , 22 Nov 1992, G. and T. Williams (1 ♀, ANIC) ; Mount Kaputar National Park: Euglah Springs road, 19 Jan 1978, E.I. Schlinger (1 ♂, CAS) ; Mount Tomah in Blue Mountains , 10 Nov 1982, 20 Nov 1982, 5 Jan 1983, and 15 Jan 1978, N.W. Rodd (4 ♂, AMS) ; 4 km W Sunny Corner at 33°22.7ʹS 149°51.6ʹE, 10 Dec 2009, V. Ahrens and W.J. Pulawski (1 ♂, CAS) , Whiskers 7 km WNW Hoskinstown at 35°24ʹS 149°23ʹE, 29 Jan 1993, M.S. Upton (1 ♀, ANIC) . Queensland: Brisbane Forest Park at 27°25ʹS 152°50ʹE, 14-29 Nov 1995, M.E. Irwin (1 ♀, MNKB) , Coast Range near Biggenden , 30 Oct 1976, H. Frauca (1 ♂, ANIC) ; Eungella National Park at 21°10.5ʹS 148°30.3ʹE, V. Ahrens and W.J. Pulawski, 31 Oct 2006 (1 ♂, CAS) , 5 Nov 2012 (1 ♂, CAS) , 7 Nov 2012 (2 ♂, CAS) , 8 Nov 2012 (1 ♂, CAS) , 9 Nov 2012 (1 ♀, CAS) . Victoria: no specific locality, date, or collector (1 ♀, BMNH) .