Pison quinquecarinatum Pulawski, 2018

Pulawski, Wojciech J., 2018, A Revision of the Wasp Genus Pison Jurine, 1808 of Australia and New Zealand, New Guinea, and the Pacific Islands (Hymenoptera: Crabronidae), Proceedings of the California Academy of Sciences 65, pp. 1-584 : 384-386

Pison quinquecarinatum Pulawski , species nova

Figures 929 -935.

NAME DERIVATION.– Quinquecarinatum is derived from the Latin numeral quinque, meaning five, and the neuter adjective carinatum , derived from carina, and meaning with carina or carinae; with reference to the presence of five longitudinal carinae on the propodeum.

RECOGNITION.– Pison quinquecarinatum is unique in having two pairs of carinae on the propodeum in addition to the median carina: the outer pair consist of the usual carinae that extend between the gastropropodeal articulation and the spiracle, and the inner pair delimit the enclosure ( Fig. 931 ). The presence of only two submarginal cells is a subsidiary recognition feature.

DESCRIPTION.– Frons dull, minutely punctate, punctures less than one diameter apart, middle supraantennal carina absent. Hypostomal carina somewhat expanded ventrally. Gena narrow in dorsal view. Labrum emarginate. Anteromedian pronotal pit either absent or transversely elongate, slightly longer than midocellar diameter. Scutum foveate along flange, with well-defined longitudinal ridges adjacent to posterior margin; scutal punctures fine, less than one diameter apart in most specimens, but about one diameter apart near center in one male; interspaces dull in female, shiny in male. Scutellum with foveate sulcus along anterior margin. Tegula enlarged, posterolaterally either impunctate or microscopically punctate, fully concealing humeral plate. Mesopleural punctures less than one diameter apart in female, in male varying from about one to about two diameters apart. Postspiracular carina absent in female, present in male (about 1.5 × as long as midocellar diameter); integument depressed between postspiracular carina and episternal sulcus. Metapleural sulcus impressed along entire length. Propodeum with irregular longitudinal carina extending from gastral socket area toward spiracle; dorsum with a pair of carinae delimiting enclosure, obliquely ridged ( Fig. 931 ), punctate between ridges (ridges inconspicuous in female, conspicuous in male); side punctate mesally, ridged along margins; posterior surface conspicuously, transversely ridged. Forewing with two submarginal cells, length of posterior margin of second cell 1.0-1.2 × its height. Hindcoxal dorsum with outer margin obtusely carinate. Punctures of tergum I about one diameter apart in female, up to about two diameters in male. Sterna finely punctate throughout.

Setae silvery, appressed on frons, gena (erect, shorter than midocellar diameter on lower gena next to occipital carina), thorax, and tergum I; nearly completely concealing integument on clypeus in female, not concealing in male). Apical depressions of terga with silvery, setal fasciae.

Head, thorax, propodeum, and gaster black, antenna black in female, ferruginous in male, mandible ferruginous, dark apically. Legs black in female, in male femora black, tibiae, and tarsi yellowish brown (tibiae black dorsally in most specimens). Mid- and hindtibial spurs whitish.

♀.– Upper interocular distance equal to 0.78 × lower interocular distance; ocellocular distance equal to 1.0 × hindocellar diameter, distance between hindocelli equal to 1.4 × hindocellar diameter; eye height equal to 0.90 × distance between eye notches. Free margin of clypeal lamella obtusely tridentate ( Fig. 929 ). Dorsal length of flagellomere I 1.4 × apical width, of flagellomere IX 0.9

× apical width. Mandible: trimmal carina with minute incision shortly beyond midlength. Length 6.2 mm; head width 1.5 mm.

♂.– Upper interocular distance equal to 0.98-1.00 × lower interocular distance; ocellocular distance equal to 1.1-1.3 × hindocellar diameter, distance between hindocelli equal to 1.2-1.3 × hindocellar diameter; eye height equal to 1.02-1.04 × distance between eye notches. Free margin of clypeal lamella acutely angulate ( Fig. 930 ). Dorsal length of flagellomere I 1.6-1.7 × apical width, of flagellomere X 0.9 × apical width. Apical margin of sternum VIII broadly, shallowly emarginate ( Fig. 932 ). Genitalia: Figs. 933, 934 . Length 4.1-4.5 mm; head width 1.2-1.3 mm.

GEOGRAPHIC DISTRIBUTION (Fig. 935).– Northern parts of Northern Territory, of Queensland, and of Western Australia.

RECORDS.– HOLOTYPE: ♂, AUSTRALIA:

Queensland: 3 km W Batavia Dawns at 12°40ʹS

142°39ʹE, 24 Oct – 23 Nov 1992, P. Zborowski and

A. Calder (ANIC).

PARATYPES: AUSTRALIA: Northern Territory :

Gregory National Park at 16°06.6ʹS 130°25.7ʹE,

12-16 June 2001, M.E. Irwin, F.D. Parker, and

C. Lambkin (1 ♂, ANIC); Kakadu National Park :

Deaf Adder Valley: Leichardt Gallery, 27 Mar 1980,

I.D. Naumann (1 ♀, ANIC). Queensland: same data as holotype (1 ♂, CAS); 7 km S Batavia Downs at

12°43ʹS 142°42ʹE, 24 Oct – 23 Nov 1992,

P. Zborowski and A. Calder (1 ♂, CAS) and 23 Nov

– 11 Dec 1992, P. Zborowski and E. Dressler (1 ♂,

ANIC). Western Australia: Lone Dingo on FIGURE 935. Collecting localities of Pison quinquecari- Mitchell Plateau at 14°35ʹS 125°45ʹE, 9-19 May natum Pulawski, sp. nov.

1983, I.D. Naumann and J.C. Cardale (1 ♀, ANIC) .

Figures

FIGURES 929-934. Pison quinquecarinatum Pulawski, sp. nov. (929) Female clypeus and mandible; (930) Male clypeus; (931) Male propodeum in lateral oblique view (arrow shows additional carina delimiting propodeal enclosure); male: (932) Sternum VIII (ventral view); (933) Genitalia in dorsal view; (934) Genitalia in lateral view.