Pison punctifemur Pulawski, 2018

Pulawski, Wojciech J., 2018, A Revision of the Wasp Genus Pison Jurine, 1808 of Australia and New Zealand, New Guinea, and the Pacific Islands (Hymenoptera: Crabronidae), Proceedings of the California Academy of Sciences 65, pp. 1-584 : 371-373

Pison punctifemur Pulawski , species nova

Figures 900 -907.

NAME DERIVATION.– Punctifemur is derived from two Latin words: punctum, a puncture, and femur, a femur; a noun in apposition to the generic name; with reference to the conspicuously punctate forefemoral venter.

RECOGNITION.– Pison punctifemur has a black gaster (with brown apical depressions of terga and golden setal fasciae), three submarginal cells, the second recurrent vein interstitial with the second intersubmarginal vein or nearly so, and setae appressed on tergum I. It has a unique combination of the conspicuous punctures on the upper frons, many of which are 2-3 diameters apart ( Fig. 902 ) and of the conspicuously large punctures on the posteroventral surface of the forefemur ( Fig. 903 ); the femoral punctures are shared with P. hirsutum . Unlike P. hirsutum , the lower gena of P. punctifemur is punctate and setose adjacent to the oral fossa (rather than nearly impunctate and asetose), the scutal punctures are no more than one diameter apart, with shiny interspaces (rather than about 2-3 diameters apart, with markedly dull interspaces), and the mesopleural punctures are less than one diameter apart (rather then more than one diameter apart near the center); also, the tergal setae of P. hirsutum are silvery.

DESCRIPTION.– Frons dull, punctate, many punctures of upper frons 2-3 diameters apart. Labrum minimally emarginate. Anteromedian pronotal pit transversely elongate, about as long as 1.5 × midocellar diameter. Scutum not foveate along flange, without short longitudinal ridges adjacent to posterior margin; scutal punctures well defined, most punctures about one diameter apart, other punctures less than one diameter apart; interspaces unsculptured. Tegula slightly enlarged. Mesopleural punctures larger than those on scutum, less than one diameter apart, up to about one diameter apart ventrally; interspaces aciculate at least in ventral half. Postspiracular carina present, about as long as midocellar diameter. Metapleural sulcus costulate between dorsal and ventral metapleural pits. Propodeum with irregular longitudinal carina separating side from dorsum and posterior surface and extending from gastral socket area toward spiracle; dorsum obliquely ridged, with or without median shallow sulcus; side ridged, punctate between ridges; posterior surface ridged, punctate between ridges. Posteroventral forefemoral surface with conspicuous punctures up to several diameters apart. Hindcoxal dorsum with outer margin not carinate. Punctures of tergum I about 1-2 diameters apart on horizontal part (less than one diameter apart adjacent to apical depression). Sternum II punctate throughout.

Setae silvery, erect on upper frons, gena (slightly sinuous, almost straight), thorax, forecoxal venter, and femoral venters; appressed on tergum I; setal length 1.5-1.0 × midocellar diameter on upper frons, 0.5-1.0 × on scutum, 2.0 × on gena; not concealing integument on clypeus in female, concealing in male. Apical depressions of terga with golden setal fasciae.

Body black, mandible all black or brown mesally, apical depressions of terga brown, tibiae ferruginous basally in some males.

♀.– Upper interocular distance equal to 0.72-0.74 × lower interocular distance; ocellocular distance equal to 1.1-1.3 × hindocellar diameter, distance between hindocelli equal to 1.1 × hindocellar diameter; eye height equal to 0.92-1.04 × distance between eye notches. Free margin of clypeal lamella arcuate ( Fig. 900 ). Dorsal length of flagellomere I 2.5-2.6 × apical width, of flagellomere IX 1.2-1.3 × apical width. Mandible: trimmal carina with small incision shortly beyond midlength. Length 10.2-10.5 mm; head width 2.9-3.1 mm.

♂.– Upper interocular distance equal to 0.86-0.90 × lower interocular distance; ocellocular distance equal to 1.8-2.0 × hindocellar diameter, distance between hindocelli equal to 1.2-1.5 × hindocellar diameter; eye height equal to 0.96 × distance between eye notches. Free margin of clypeal lamella acutely angulate, its lateral margin slightly concave in most specimens ( Fig. 901 ). Dorsal length of flagellomere I 2.4-2.8 × apical width, of flagellomere X 1.1-1.2 × apical width.

Sternum VIII emarginate apically ( Fig. 904 ). Genitalia: Figs. 905, 906 . Length 8.9-11.2 mm; head width 2.7-3.2 mm.

GEOGRAPHIC DISTRIBUTION (Fig. 907).– Western Australia.

RECORDS.– HOLOTYPE: ♀, AUSTRALIA: Western Australia: 60 km NE Wubin at 29°43ʹS 117°04ʹE, 27 Sept 1981, I.D. Naumann and J.C. Cardale ( ANIC).

PARATYPES: AUSTRALIA: Western Australia: Mount Gibson Station at 29°36ʹ52ʺS 117°24ʹ38ʺE, 25 Sept 2001, S.R. Paterson (1 ♂, WAM) ; Mount Gibson Station 93 km NE Wubin at 29°41.23ʹS 117°21.62ʹE, 21-29 Aug 2001, R. Leys and K. Ottewell (2 ♀, SAM) ; same data as holotype (5 ♀, 18 ♂, ANIC; 3 ♀, 9 ♂, CAS) .

Figures

FIGURES 900-903. Pison punctifemur Pulawski, sp. nov. (900) Female clypeus; (901) Male clypeus; (902) Female frons; (903) Female forefemur (posteroventral surface).

FIGURES 904-906. Pison punctifemur Pulawski, sp. nov., male. (904) Sternum VIII (ventral surface); (905) Genitalia in dorsal view; (906) Genitalia in lateral view. FIGURE 907. Collecting localities of Pison punctifemur Pulawski, sp. nov.