Pison oculare Pulawski, 2018

Pulawski, Wojciech J., 2018, A Revision of the Wasp Genus Pison Jurine, 1808 of Australia and New Zealand, New Guinea, and the Pacific Islands (Hymenoptera: Crabronidae), Proceedings of the California Academy of Sciences 65, pp. 1-584 : 315-317

Pison oculare Pulawski , species nova

Figures 749 -753.

NAME DERIVATION.– Oculare is the Latin neuter adjective meaning ocular; with reference to the shallowly emarginate eyes of this species.

RECOGNITION.– Like Pison orbitale , P. oculare has an unusually shallow eye emargination, whose depth is less than half of midocellar diameter ( Fig. 750 ), whereas in all other Pison the emargination is about as deep as the midocellar diameter. Both species also have a fine omalus, which is found only exceptionally in the other Pison (e.g., P. tenebrosum ). Additionally, the second recurrent vein is received near the midlength of the second submarginal cell ( Figs. 751, 752 ). See Pison orbitale for differences between the two species (p. 317).

DESCRIPTION.– Frons dull, minutely punctate, punctures less than one diameter apart. Occipital carina narrowly separated from hypostomal carina. Gena narrow in dorsal view. Eye emargination unusually shallow, less than half as deep as midocellar diameter ( Fig. 750 ). Labrum not emarginate. Anteromedian pronotal pit transversely elongate, about as long as 1.5 × midocellar diameter. Scutum foveate along flange, without longitudinal ridges adjacent to posterior margin; scutal punctures minute, interspaces linear, microsculptured, dull. Tegula enlarged. Mesopleuron with fine omalus (closer to anterior margin than in orbitale ); mesopleural punctures markedly larger than those on scutum, interspaces linear. Postspiracular carina present, about 1.5 × as long as midocellar diameter; integument depressed between postocellar carina and episternal sulcus. Metapleural sulcus not costulate between dorsal and ventral metapleural pits. Propodeum with irregular longitudinal carina separating side from dorsum and posterior surface and extending from gastral socket area toward spiracle (carina obscured dorsally by adjacent rugae); dorsum conspicuously rugose, with middle carina but without sulcus; side ridged, punctate between ridges; posterior surface irregularly rugose (specimen from Wilpena) or irregularly ridged (specimen from Taree area), with several conspicuous ridges radiating up from transverse carina just above gastropropodeal articulation. Forewing with three submarginal cells ( Figs. 751, 752 ), submarginal cell II greatly reduced in size in specimen from Taree area ( Fig. 752 ), its height about 0.2 of distance that separates it from marginal cell; second recurrent vein ending near middle of submarginal cell II.

ter; eye height equal to 1.14-1.16 × distance between eye notches. Clypeus mesally with well-defined punctures, many of which are more than one diameter apart; free margin of lamella slightly, evenly arcuate, rounded laterally ( Fig. 749 ). Dorsal length of flagellomere I 2.1-2.3 × apical width, of flagellomere IX 1.5. × apical width. Mandible: trimmal carina without incision; acetabular groove with two rows of punctures. Length 4.6-5.9 mm; head width 1.6-1.7 mm.

♂.– Unknown.

GEOGRAPHIC DISTRIBUTION (Fig.753).– New South Wales, South Australia.

RECORDS.– HOLOTYPE: ♀, AUSTRALIA: New South Wales: Doyles River 50 km NW Taree at 31°31ʹS 152°14ʹE, 15 Nov 2009, D. Bray ( AMS).

PARATYPE: AUSTRALIA: South Australia: Wilpena in Flinders Ranges National Park at 31°31.7ʹS 138°36.2ʹE, 22 Dec 2010, V. Ahrens and W.J. Pulawski (1 ♀, CAS) .

Figures

FIGURES 749-752. Pison oculare Pulawski, sp. nov., female. (749) Clypeus and mandibles; (750) Head in frontal view; (751) Portion of left forewing (arrow shows second submarginal cell); (752) Portion of right forewing (arrow shows second submarginal cell). FIGURE 753. Collecting localities of Pison oculare Pulawski, sp. nov.