Pison terrigena Pulawski, 2018

Pulawski, Wojciech J., 2018, A Revision of the Wasp Genus Pison Jurine, 1808 of Australia and New Zealand, New Guinea, and the Pacific Islands (Hymenoptera: Crabronidae), Proceedings of the California Academy of Sciences 65, pp. 1-584 : 455-457

Pison terrigena Pulawski , species nova.

Figures 1108-1113.

NAME DERIVATION.– Terrigena is a Latin noun meaning born from the earth, son of the earth, a noun in apposition to the generic name; also with reference to this species origin in the Northern Territory of Australia.

RECOGNITION.– The male of P. terrigena (the female is unknown) is all black (mandible yellowish) and has three submarginal cells, the propodeum without a longitudinal carina separating the side from the dorsum and the posterior surface, the second recurrent vein interstitial with the second intersubmarginal vein or nearly so, and the setae appressed on tergum I. It differs from similar species by the following combination: punctures of frons and scutum well defined; clypeal lamella acutely angulate; scutal punctures less than one diameter apart; ocellocular distance equal to 0.8 × hindocellar diameter; dorsal length of flagellomere I 2.0 × apical width; setae of lower gena nearly as long as midocellar diameter; sternum II with large punctures, impunctate apicomesally; sterna IV and V with a few sparse punctures (except punctures closer to each other near the lateral margin); apical margin of sternum VIII slightly emarginate and with obtuse apicolateral corner (Fig. 1110).

DESCRIPTION.– Frons dull, finely punctate, punctures somewhat ill defined, less than one diameter apart, supraantennal carina replaced by thin sulcus. Occipital carina joining hypostomal carina. Gena narrow in dorsal view (Fig. 1109). Labrum not emarginate. Anteromedian pronotal pit transversely elongate, slightly longer than midocellar diameter. Scutum not foveate along flange, without longitudinal ridges adjacent to posterior margin; scutal punctures well defined, nearly contiguous except narrowly separated laterally in holotype (less than one diameter apart). Tegula not enlarged. Mesopleural punctures contiguous. Postspiracular carina present, almost as long as midocellar diameter. Metapleural sulcus costulate between dorsal and ventral metapleural pits. Propodeum without longitudinal carina separating side from dorsum and posterior surface; dorsum rugose, with short transverse ridges emerging from midline; side finely ridged, rugose between ridges; posterior surface inconspicuously, irregularly ridged transversally, rugose between ridges. Posteroventral forefemoral surface minutely, closely punctate. Punctures of tergum I, anterior of apical depression, averaging about one diameter apart (except nearly contiguous laterally). Sternum II with large punctures, impunctate apicomesally; sterna IV and V with a few sparse punctures (except punctures closer to each other near lateral margin).

Setae silvery, appressed on frons, postocellar area, scutum, and tergum I; oriented dorsally on upper frons (except near orbit); on lower gena curved, nearly as long as midocellar diameter, mostly subappressed, but a few setae erect; completely concealing integument on clypeus (except lamella). Apical depressions of terga with silvery, setal fasciae.

Body black, mandible yellowish, brown apically.

♀.– Unknown.

♂.– Upper interocular distance equal to 0.76 × lower interocular distance; ocellocular distance equal to 0.8 × hindocellar diameter, distance between hindocelli equal to 0.9-1.0 × hindocellar diameter; eye height equal to 0.96-1.00 × distance between eye notches. Free margin of clypeal lamella acutely angulate (Fig. 1109). Dorsal length of flagellomere I 2.0 × apical width, of flagellomere X 1.2 × apical width. Sternum VIII with apical margin shallowly emarginate, with obtuse apicolateral corner (Fig. 1110). Genitalia: Figs. 1111, 1112. Length 6.5-7.5 mm; head width 1.9-2.1 mm.

GEOGRAPHIC DISTRIBUTION (Fig. 1113).– Northern part of Northern Territory.

RECORDS.– HOLOTYPE: ♂, AUSTRALIA: Northern Territory: Keep River National Park at 15°57ʹ55ʺS 129°01ʹ52ʺE, 10-13 June 2001, M.E. Irwin, F.D. Parker, and C. Lambkin ( ANIC).

PARATYPE: AUSTRALIA: Northern Territory: Gregory National Park at 16°12ʹ47ʺS 130°25ʹ11ʺE ,

12-15 June 2001, M.E. Irwin, F.D. Parker, and C. Lambkin (1 ♂, CAS) .