Pison trilobatum Pulawski, 2018

Pulawski, Wojciech J., 2018, A Revision of the Wasp Genus Pison Jurine, 1808 of Australia and New Zealand, New Guinea, and the Pacific Islands (Hymenoptera: Crabronidae), Proceedings of the California Academy of Sciences 65, pp. 1-584 : 473-474

Pison trilobatum Pulawski , species nova

Figures 1149-1151.

NAME DERIVATION.– The name trilobatum derives from the Latin prefix tri-, meaning three, and the adjective lobatus (neuter: lobatum), meaning having lobes, lobate; with reference to the trilobate clypeal lamella of this species female.

RECOGNITION.– Pison trilobatum has only two submarginal cells, an entirely, finely punctate tegula, and, in the female (the male is unknown), the dorsal length of flagellomere II is 2.4 × its apical width. Pison incurvatum and P. bicellulare are similar, but in P. trilobatum the legs are ferruginous (rather than black), the clypeal lamella is distinctly trilobate (Fig. 1149), not bent posterad (only slightly trilobate in bicellulare , evenly arcuate and bent posterad in P. incurvatum ).

DESCRIPTION.– Frons dull, minutely punctate, punctures almost contiguous. Distance between antennal socket and orbit slightly smaller than socket width. Gena narrow in dorsal view (Fig. 1150). Labrum not emarginate. Anteromedian pronotal pit rounded, slightly transversely elongate, slightly shorter than midocellar diameter. Scutum slightly foveate along flange, with short, evanescent longitudinal ridges adjacent to posterior margin; scutal punctures minute, less than one diameter apart. Scutellum with foveate sulcus along anterior margin. Tegula enlarged. Mesopleural punctures fine, about one diameter apart. Postspiracular carina present, about half as long as midocellar diameter. Metapleuron microscopically punctate; metapleural sulcus not costulate between dorsal and ventral metapleural pits. Propodeum with irregular longitudinal carina separating side from dorsum and posterior surface and extending from gastral socket area toward spiracle; dorsum obliquely (almost transversely) ridged; side finely ridged, punctate between ridges (impunctate anteriorly), punctures mostly more than one diameter apart, but less than that posteriorly; posteri- or surface transversely ridged, punctate between ridges. Forewing with two submarginal cells; posterior margin of second submarginal cell equal to 1.1 × its height. Hindcoxal dorsum with outer margin obtusely carinate. Outer surface of hindtibia with evanescent spines. Punctures of tergum I minute, averaging slightly more than one diameter apart on horizontal portion. Sterna minutely punctate throughout.

Setae silvery, appressed on frons, postocellar area, gena, scutum, and tergum I; not concealing integument on clypeus. Apical depressions of terga I and II with inconspicuous, silvery, setal fasciae; following terga with evanescent fasciae.

Head, thorax, propodeum, and gaster black, lamella of female clypeus and adjacent portion of clypeus proper ferruginous; mandible yellowish (brown at very apex); labrum yellow; antenna ferruginous ventrally, black dorsally. Trochanters, femora, tibiae, and tarsi ferruginous; mid- and hindtibial spurs whitish.

♀.– Upper interocular distance equal to 1.0 × lower interocular distance; ocellocular distance equal to 0.5 × hindocellar diameter, distance between hindocelli equal to 1.1 × hindocellar diameter; eye height equal to 1.06 × distance between eye notches. Free margin of clypeal lamella distinctly trilobate, median lobe widest (Fig. 1149). Dorsal length of flagellomere I 2.4 × apical width, of flagellomere IX 1.1 × apical width.

Mandible: trimmal carina with minuscule incision at about apical two thirds of length.

Length 4.3 mm; head width 1.0 mm.

♂.– Unknown.

GEOGRAPHIC DISTRIBUTION Fig. 1151).–

Known from two closely adjacent localities in

Western Australia.

RECORDS.– HOLOTYPE: ♀, AUSTRALIA: Western Australia : Karijini National Park at 22°25.6ʹS

118°23.7ʹE, 23 Apr – 4 May 2003, F.D. Parker and

M.E. Irwin (ANIC).

PARATYPE: AUSTRALIA: Western Australia :

Karijini National Park at 22°26.3ʹS 118°22.9ʹE,

23 Apr – 4 May 2003, F.D. Parker and M.E. Irwin

FIGURE 1151. Collecting localities of Pison trilobatum Pulawski , sp. nov.

( 1 ♀, CAS) .