Pison westwoodii Shuckard

Pulawski, Wojciech J., 2018, A Revision of the Wasp Genus Pison Jurine, 1808 of Australia and New Zealand, New Guinea, and the Pacific Islands (Hymenoptera: Crabronidae), Proceedings of the California Academy of Sciences 65, pp. 1-584 : 487-498

Pison westwoodii Shuckard

Figures 1183-1197.

Pison westwoodii Shuckard, 1838:77 , ♀ (as Westwoodii , incorrect original capitalization). Holotype or syntypes, ♀, Australia: Van Diemen’s Land, now Tasmania: no specific locality (lost). Neotype: ♀, lectotype of Pison obliquum (OXUM) , present designation. – F. Smith, 1956:316 (in catalog of Hymenoptera in British Museum), 1869:290 (in checklist of Pison ); Kohl, 1885:189 (in checklist of world Pison ); Froggatt, 1892:218 (in catalog of Australian Hymenoptera ); Dalla Torre, 1897:713 (in catalog of world Hymenoptera ); Turner, 1915:558 (in key to Pison of Tasmania, relationship to Pison iridipenne ), 1916b:597 (in key to Australian Pison ), 604 (variation, recognition characters, Pison obliquum is probably a synonym); R. Bohart and Menke, 1976:337 (in checklist of world Sphecidae ); Evans, Matthews, and Hook, 1981:224 (nesting behavior); Cardale, 1985:263 (in catalog of Australian Sphecidae ); Naumann, 1990a:24 (Norfolk and Philips Islands); K. Walker, Naumann, Austin, Taylor, and Cardale, 1992:49 (in catalog of insects of Tasmania); Naumann, 1993:185 ( Australia: Queensland: Heathlands area in Cape York); Baker, 1998:173 (origin and depository of type material); Smithers, 1998:46 (in list of insects of Norfolk Island).

Pison obliquum F. Smith, 1856:316 , ♀ (as obliquus, incorrect original termination). Lectotype: ♀, Australia: Van Diemen’s Land (now Tasmania): no specific locality (OXUM), p resent designation, examined. New synonym. – F. Smith, 1869:291 (in checklist of Pison ); Turner, 1916b:604 (a tentative synonym of Pison westwoodii ). – As Pison obliquum : Kohl, 1885:187 (in checklist of world Pison ); Froggatt, 1892:217 (in catalog of Australian Hymenoptera ); Dalla Torre, 1897:712 (in catalog of world Hymenoptera ); R. Bohart and Menke, 1976:337 (as questionable synonym of Pison westwoodii ); Cardale, 1985:263 (in catalog of Australian Sphecidae , as junior synonym of Pison westwoodii ).

Pison iridipenne F. Smith, 1879a:676 , ♀, ♂ (as iridipennis , incorrect original termination). Lectotype: ♂, Hawaiian Islands: Oahu: Honolulu (BMNH), present designation, examined. New synonym. – Blackburn and Kirby, 1880:88 (Hawaiian Islands); Kohl, 1885:187 (in checklist of world Pison ); Blackburn and Cameron, 1886:173 (Hawaiian Islands: Honolulu); Dalla Torre, 1897:711 (in catalog of world Hymenoptera ); R. Perkins in R. Perkins and Forel, 1899:14 (Hawaiian Islands: Honolulu), 1901:264 (Hawaiian Islands); Turner, 1908:512 Queensland: Cairns, Mackay), 1916b:626 (bibliographic references); Bridwell, 1919b:123 (in key to Hawaiian Pison ); Giffard, 1919:181 ( American Samoa and Hawaii); F. Williams, 1927:438 (common about Honolulu); Cheesman, 1928:176 (Marquesas and Society Islands); Perkins and Cheesman, 1928:6 (listed from Samoa), 28 ( Samoa, distinctive characters); F. Williams, 1932 (Marquesas Islands); Krauss, 1944:93 (Hawaii: Molokai Island); F. Williams, 1947:318 and 331 (not known from Fiji); Krombein, 1949b:386 (in key to Sphecidae of Micronesia), 408 (geographic variation; Marshall, Mariana, and Caroline Islands), 1950:139 (additional Micronesian localities); Yasumatsu, 1953:141 (in list of Pison of Pacific islands), 141 (bibliographic references; Micronesia); Fullaway, 1957:279 (in checklist of Hymenoptera of Fiji); Yoshimoto, 1960:334 (in list of Hawaiian Sphecidae ); Baltazar, 1966:335 (in catalog of Hymenoptera of Philippines); Hinckley 1969:15 ( Tokelau Islands); R. Bohart and Menke, 1976:336 (in checklist of world Sphecidae ); Tsuneki, 1976:94 ( Philippines; redescription); Menke, 1979:303 (Tahiti); Tsuneki, 1982a:38 ( Bismarck Archipelago, redescription), 1983b:42 (in key to Pison of New Guinea), 43 (New Guinea); Cardale, 1985:260 (in catalog of Australian Sphecidae ); Menke, 1990:154 ( doggonum may be a junior synonym); Evenhuis, 2007:6 (in checklist of Hymenoptera of Fiji).

Pison iridipenne F. Smith, 1879b:139 , ♀, ♂ (as iridipennis , incorrect original termination). Objective synonym of Pison iridipenne F. Smith, 1879a:676 .

Pison strictifrons Vachal, 1907:114 , ♀. Syntypes: New Caledonia: no specific locality (MNHN), photographs examined. New synonym. – R. Turner, 1916g:626 (bibliographic reference, morphological characters); Williams, 1945:440 ( New Caledonia, recognition characters); Yasumatsu, 1953:134 (in list of Pison of Pacific islands); R. Bohart and Menke, 1976:336 (in checklist of world Sphecidae ); Callan, 1990:20 ( New Caledonia: no specific locality); Rasmussen, 2012:46 (in list of species described by Vachal); Jennings, Krogmann, and Burwell, 2013:32 (in checklist of Hymenoptera of New Caledonia).

Pison impunctatum Turner, 1912a:200 , ♀. Holotype by monotypy: ♀, Indonesia: Western Papua (= Indonesian New Guinea): Mimika River (BMNH), examined. New synonym. – Turner in Turner , Meade-Waldo, and Morley, 1915:6 (New Guinea: Mimika River ; redescription); Turner, 1916b:626 (recognition characters, New Guinea); Cheesman, 1928:176 (Marquesas and Society Islands , description of ♂); R. Bohart and Menke, 1976:336 (in checklist of world Sphecidae ); Tsuneki, 1983b:42 (in key to Pison of New Guinea).

Pison korrorense Yasumatsu, 1937b:133 , ♀, ♂. Holotype: ♀, Palau Republic: Island of Koror (ELKU), examined. New synonym. – Yasumatsu, 1939b:82, 83 (in key to Pison of eastern Asia, in checklist of Pison of Japanese Empire); Krombein, 1949b:385 (in key to Sphecidae of Micronesia, description of ♂), 409 (possibly a subspecies of iridipenne , Caroline Islands), 1950b:134 and 139 (illustrations of head, sternum III, and genitalia); Yasumatsu, 1953:134 (in list of Pison of Pacific islands), 141 (bibliographic references; Micronesia); R. Bohart and Menke, 1976:336 (in checklist of world Sphecidae ; possibly synonym of Pison iridipenne ).

Pison doggonum Menke, 1988a:26 , ♀. Holotype: ♀, Mexico: no specific locality, corrected to Western Pacific Islands by Menke , 1990:154, may actually be southeast Asia , New Guinea, or Australia (ZMUC), examined. New synonym. – Menke, 1990:154 (possibly a synonym of iridipenne F. Smith ).

NEOTYPE AND LECTOTYPE DESIGNATIONS.– According to the original description (Shuckard, 1838), the type(s) of Pison westwoodii belonged to J.O. Westwood, whose collection is now preserved in the Hope Entomological Collections, Oxford. Unfortunately, no specimen of this species can be found there, as James E. Hogan, responsible for the Hymenoptera , informed me in his e-mails of 6 May and 15 July 2011. Shuckard’s type material is not present in the Natural History Museum, London, either (information of David G. Notton). I accept therefore that the original type material has been lost. In order to fix this common species interpretation, I hereby select as the neotype the lectotype of Pison obliquum F. Smith, 1856 (see next paragraph). Both taxa originat- ed from Van Diemen’s Land (now Tasmania), with no specific locality. The original description of westwoodii does not allow an unequivocal recognition, but the neotype clearly agrees with the interpretation of Turner, 1916b, the only previous reviser of the Australian Pison .

Smith (1856) did not indicate the number of specimens examined in the original descriptions of Pison obliquum , but he gave Van Diemen’s Land (now Tasmania) as the country of origin and the collection of W.W. Saunders as depository. Two specimens of this species are present in the Hope Entomological Collections, Oxford, under a handwritten label “ obliquus, V.D.L”. I have selected as the lectotype the one bearing a square handwritten label “V.D.L.”, and the other one with undecipherable letters on a square label as the paralectotype.

Pison iridipenne F. Smith, 1856 , was described from both sexes from Honolulu, Hawaii. I have selected as the lectotype the only existing original specimen in the Natural History Museum, London.

RECOGNITION.– Pison infumatum , P. nitens , and P. westwoodii are all black, have three submarginal cells, the second recurrent vein contiguous with the second intersubmarginal vein or nearly so, and the setae appressed on tergum I. The females are unique among the Australian species in having the ocellocular distance equal to 0.1-0.5 × hindocellar diameter (Fig. 1188) in combination with a sparsely punctate mesopleuron (Fig. 1190), with punctures slightly more than one diameter apart to several diameters apart. They differ as follows: in P. westwoodii , the frontal punctures are more than one diameter apart, the propodeum of most specimens has a longitudinal carina separating the dorsum and posterior surface from the side and extending from the gastral socket area toward the spiracle, and the propodeal dorsum of most specimens is at least slightly ridged, with most or all punctures no more than one diameter apart (only the punctures are present except the basal ridges and the median sulcus in some specimens). In the other two species, the frontal punctures are about one diameter apart, there is no longitudinal carina separating the propodeal dorsum and posterior surface from the side, the propodeal dorsum is punctate only, except ridged next to anterior margin, with punctures more than one diameter apart.

The vast majority of males of P. westwoodii share with P. morosum (an endemic. of New Zealand, where P. westwoodii does not occur) a sparsely punctate, transverse swelling on sternum III, slightly behind its midlength (Fig. 1191, 1192); in some specimens a swelling is also present on sternum IV. The swelling is entire in most specimens, separated mesally into two sections in some (Fig. 1192). The two species can be differentiated by the following: in P. westwoodii , the setae of the upper frons are silvery, 0.3-0.4 × as long as midocellar diameter just below the midocellus, the apical portion of sternum II is microscopically punctate, and sternum II is simple. In P. morosum , the setae of upper frons are brown and up to 0.7 × as long as the the midocellar diameter just below the midocellus, the apical portion of sternum II is impunctate, and sternum II has an ill-defined, preapical, medially divided tranverse swelling. Somewhat similar are certain males of P. sulcatum , in which sternum IV in many specimens (also sternum III in many specimens) has a pair of sharp tubercles (Fig. 1076). Sternum VIII of P. westwoodii , however, is simple, whereas in P. sulcatum it has a longitudinal sulcus or a round concavity (Fig. 1077-1080).

All known males from New Caledonia and some rare males from other areas lack the swelling on sternum II. Such specimens can be recognized by the relatively sparse mesopleural punctures that are slightly more to markedly more than one diameter apart, and the subsidiary recognition features are: body all black, ocellocular distance equal to 0.3-0.6 × hindocellar diameter, sterna punctate throughout, and sternum VIII moderately emarginate apically (Fig. 1193). Pison modestum is similar, but in that species the posterior propodeal surface is punctate in at least the dorsal half (posterior propodeal surface all ridged in P. westwoodii ).

JUSTIFICATION OF NEW SYNONYMY.– The syntypes of P. obliquum and the holotypes of P. impunctatum and P. doggonum are clearly conspecific with P. westwoodii , and I treat them as junior synonyms of the latter name. I regard a different shape of the clypeal lamella in the Hawaiian males (described as P. iridipenne ) as the result of geographic variation, and I synonymize this name with P. westwoodii , too.

I could not examine the syntypes of Pison strictifrons , but I have seen the photographs of one female with the original handwritten determination label that Mademoiselle Agnièle Touret-Alby (Muséum National d’Histoire Naturelle, Paris, France) sent me on 1 August 2017. The photographs clearly show the ocellocular distance smaller than the hindocellus diameter, short setae of the lower gena, the mesopleural punctures more than one diameter apart, the second recurrent vein interstitial with the second intersubmarginal vein, and the propodeal dorsum obliquely ridged. Of the five species currently known to occur in Nova Caledonia , P. argentatum , P. marginatum , P. novocaledonicum , P. rufipes , and P. westwoodii , the first four are clearly different from P. strictifrons , which, however, shows all the characters of P. westwoodii . Therefore I synonymize these two names.

The female holotype of P. korrorense is identical to specimens of P. westwoodii from Australia. The only known male, described by Krombein (1949), resembles the Australian P. westwoodii except for the punctures of the upper frons that are nearly contiguous. I regard it as P. westwoodii because of the presence of a sparsely punctate, transverse swelling on its sternum III.

DESCRIPTION.– Frons bulging above antennal sockets, dull, minutely punctate, punctures more than one diameter apart in female, varying from more than to less than one diameter apart in male. Labrum emarginate. Anteromedian pronotal pit transversely elongate, about as long as midocellar diameter. Propleuron either punctate throughout or impunctate laterally. Scutum not foveate or inconspicuously foveate along flange, but conspicuously foveate in some males from New Guinea, with short longitudinal ridges adjacent to posterior margin; scutal punctures minute, less than one diameter apart to more than one diameter apart (see Geographic Variation below). Tegula not enlarged. Mesopleural punctures ranging from minute, inconspicuous to well-defined (see Geographic Variation below), slightly more than one diameter apart to several diameters apart; interspaces microsculptured, dull except unsculptured in some specimens from Koror Island and some from Pohnpei Island, Pacific Ocean. Postspiracular carina present, about as long as midocellar diameter. Metapleural sulcus not costulate between dorsal and ventral metapleural pits, metapleural punctures microscopically small. Propodeum in most specimens with longitudinal carina separating side from dorsum and posterior surface and extending from gastral socket area toward spiracle (carina evanescent or absent in many specimens); dorsum with ridges varying from conspicuous to evanescent, punctate between ridges; in specimens with evanescent ridges punctures become apparent (only punctures present except basal ridges and median sulcus in some specimens); side ridged, punctate between ridges, but only punctate in many males; posterior surface ridged, punctate between ridges, conspicuously rugose in exceptional specimens. Hindcoxal dorsum with outer margin sharply carinate posteriorly. Punctures of tergum I minute, averaging slightly more than one diameter apart. Sterna punctate throughout.

Setae silvery, shorter than midocellar diameter, inclined obliquely ventrally between midocellus and dorsal end of midfrontal carina, appressed on scutum and tergum I; suberect, up to about 0.7 × midocellar diameter on lower gena; not concealing integument on clypeus in female, partly concealing in male. Apical depressions of terga with inconspicuous, silvery, setal fasciae.

Body all black except mandible dark ferruginous mesally.

♀.– Upper interocular distance equal to 0.48-0.54 × lower interocular distance; ocellocular distance equal to 0.1-0.5 × hindocellar diameter, distance between hindocelli equal to 0.4-1.1 × hindocellar diameter; eye height equal to 1.06-1.10 × distance between eye notches. Free margin of clypeal lamella rounded (Fig. 1183), obtusely angulate in some specimens. Dorsal length of flagellomere I 2.2-2.6 × apical width, of flagellomere IX 1.2-1.3 × apical width. Mandible: trimmal carina with small incision shortly beyond midlength. Length 6.5-8.1 mm; head width 1.9-2.3 mm.

♂.– Upper interocular distance equal to 0.62 × lower interocular distance; ocellocular distance equal to 0.3-0.6 × hindocellar diameter (Fig. 1189), distance between hindocelli equal to 0.7-0.9 × hindocellar diameter; eye height equal to 1.12-1.40 × distance between eye notches. Free margin of clypeal lamella acutely angulate to rounded (see Geographic Variation below). Dorsal length of flagellomere I 2.5-2.7 × apical width, of flagellomere X 1.2-1.3 × apical width. Sternum III mesally in the overwhelming majority of specimens (also sternum IV in some specimens) with sparsely punctate, transverse swelling (Fig. 1191) slightly behind midlength (see below for details); swelling divided mesally in many specimens (Fig. 1191), reduced to median tubercle in some others, forming longitudinal projection in specimens from Sulawesi, and lacking altogether in some rare specimens (see below for details). Sternum VIII broadly emarginate apically (Fig. 1193), in lateral view: Fig. 1194. Genitalia: Figs. 1195, 1196. Length 6.0- 8.3 mm; head width 1.6-2.2 mm.

MALE STERNUM III. – The vast majority of males has a transverse swelling mesally on sternum III (Figs. 1191, 1192), but the swelling lacks altogether in some specimens. These two forms are othewise identical, both externally and in the the shape of their genitalia, and some of them were collected in mixed populations. For these reasons I treat them as conspecific. The following specimens have no sternal swelling:

In Australia, one from 4 km W Orford, Tasmania.

Two of the five males from Saipan Islands, Marianas .

The only male from Vanuatu.

The only male from Singkor, Malaysia.

All males from New Caledonia.

The three males examined from Philippines .

GEOGRAPHIC VARIATION.– Singapore, Borneo: Scutal punctures more than one diameter apart; mesopleuron microsculptured, but less so than in Australian populations, punctures well defined. Male: clypeal lamella about rectangular, rounded apically.

Australia: Scutal punctures about one diameter apart in most specimens, less than one diameter apart in some; mesopleuron conspicuously microsculptured (but only slightly so in some specimens), punctures minute, ill defined in most specimens, but somewhat larger, well defined in those from Bowling Green Bay National Park, 9 and 18 km S Dingo Beach, and Split Rock , Queensland, some from Emerald and Homevale National Park, Queensland, and in those from Kalumburu Mission area, Mining Camp, and Lone Dingo, Western Australia. Male: clypeal lamella acutely angulate to rectangular (Fig. 1184).

New Guinea: Scutal punctures more than one diameter apart in most specimens, but about one diameter apart in some; mesopleuron microsculptured, but less so than in the Australian populations, punctures well defined. Male clypeal lamella obtusely angulate (Fig. 1185), prominently rounded in some specimens.

New Caledonia: Male clypeal lamella varying from acutely angulate to obtusely angulae.

American Samoa: Male clypeal lamella roundly angulate in one specimen examined .

Fiji: Scutal punctures about one diameter apart; mesopleuron slightly microsculptured, punctures well defined. Male: clypeal lamella prominently rounded (Fig. 1186).

Cooks Islands: As above, but clypeal lamella of male obtusely angulate.

Hawaiian Islands: As above, but clypeal lamella of male short, rounded (Fig. 1187).

Pitcairn Island: As above, male unknown.

NESTING HABITS.– According to Williams (1927), this species (as P. iridipennis ) “ is common about Honolulu, Hawaii and has been found nesting in the old twig tips of the Night-Blooming Cereus ” (i.e., Hylocereus undatus Britton and Rose ). Evans, Matthews, and Hook (1981) described three nests of P. westwoodii , all established in artificial nest in Canberra, A.C.T. area. The nests contained three, four and seven cell, respectively, ranging from 6.5 to 14 mm in length, separated by mud partitions 1-2 mm thick. One of the nests had an empty vestibular cell that was closed off by a thick mud plug. Another nest contained in two cells many tiny spiderlings, ranging 19 and 30 per cell. The authors also observed two specimens that emerged from a nest of P. rufipes and commented that their mother had probably appropriated cells in a nest constructed by P. rufipes , as P. westwoodii is known to occupy hollow cavities.

GEOGRAPHIC DISTRIBUTION (Fig. 1197).– Singapore, Indonesia, Thailand, Philippines, New Guinea, Australia, Solomon Islands, Caroline Islands, Mariana Islands, Marshall Islands, Fiji,

Samoa, American Samoa, French

Polynesia, Cook Islands, Hawaiian Islands, Pitcairn Islands;

from sea level up to 2,500 m in

Papua New Guinea.

RECORDS.– AMERICAN SAMOA:

Tutuila: Aunuu Island (1 ♀, BISH) ,

Fagasa Trail (1 ♀, BISH), LeoneAluau Trail (1 ♀, BISH), Pago Pago

(1 ♀, BISH), Taputimu (1 ♂, BISH) ,

Vailoatai (1 ♀, 1 ♂, BISH), Vaitogi

(1 ♀, BISH), no specific locality

(Giffard, 1919, as iridipennis ).

AUSTRALIA: Australian Capi-

FIGURE 1197. Collecting localities of Pison westwoodii Shuckard.

tal Territory: Black Mountain at

35°16ʹS 146°06ʹE (5 ♀, 1 ♂, ANIC; 2 ♀, 1 ♂, CAS), Blundells (1 ♂, ANIC) , Canberra (14 ♀, 6 ♂, ANIC) , Cotter River at Bendora Creek (4 ♀, CAS) , Tidbinbilla Nature Reserve (1 ♀, AMS) . Christmas Island: no specific locality (2 ♀, BISH) . New South Wales : Barrington House via Salisbury (1 ♀, QMB) , Boonoo Boonoo National Park (1 ♀, AMS) , Burrendong Botanic Garden at 32°42.1ʹS 149°06.2ʹE (2 ♂, CAS) , Clarence (1 ♀, 2 ♂, AMS) , 119 km W Cobar at 31°33.5ʹS 144°37.6ʹE (1 ♀, CAS) , Congo 8 km ESE Moruya at 35°58ʹS 150°09ʹE (9 ♀, ANIC) , Coocumbac Island Nature Reserve near Taree (1 ♀, ANIC) , Coolbaggie Forest Reserve 10 km E Eumungerie at 31°58.5ʹS 148°40.5ʹE (9 ♀, CAS) , Doyles River 50 km NW Taree at 31°31ʹS 152°14ʹE (3 ♀, AMS) , 1 km W Eumungerie at 31°56.7ʹS 148°36.9ʹE (5 ♀, 1 ♂, CAS) , Forbes (1 ♀, SAM) , Gibraltar Range National Park (1 ♂, ANIC) , Gilgandra Flora Reserve at 31°39.7ʹS 148°46.3ʹE (1 ♂, CAS) , 13 km NW Harden (3 ♀, 3 ♂, ANIC) , Jervis Bay : Hyam’s Beach (1 ♀, ANIC) , Kamay Botany National Park 14 km S center of Sydney at 34°00.3ʹS 151°13.2ʹE (3 ♀, CAS) , Kiandra : Alpine Creek (1 ♀, ANIC) , Kinchega National Park at 32°23.7ˈS 142°22.7ˈE (4 ♀, 7 ♂, CAS) , Kosciuszko National Park : Olgives Creek near Round Mountain (1 ♂, CAS) , Lake George Cullerin (1 ♀, 1 ♂, UCD) , Lansdowne near Taree (1 ♀, AMS) , 3 km N Lansdowne near Taree (1 ♀, ANIC) , 0.5 km SE Lansdowne near Taree (1 ♀, AMS; 1 ♀, ANIC), Lindfield at 33°46ʹS 151°11ʹE (4 ♀, 1 ♂, ANIC) , Little River in Blue Mountains (2 ♂, AMS) , Lord Howe Island : ca 500 m NW Mutton (1 ♀, AMS) , Lord Howe Island : Research Station backyard at 31°31ʹ37ʺS 159°03ʹ58ʺE (8 ♀, 5 ♂, AMS) , Lorien Wildlife Refuge 3 km N and ca 1 km NNW Lansdowne near Taree (2 ♀, AMS) , Manly : Kangaroo Park (4 ♀, ANIC) , Mount Banks in Blue Mountains (1 ♀, AMS) , Mount Kaputar National Park (1 ♀, AMS) , Mount Tomah (1 ♂, AMS) , 16 km N Mudgee (1 ♀, 5 ♂, ANIC) , Myall Lakes National Park : Mungo Brush (2 ♂, AMS) , New England National Park (1 ♀, CAS) , Nowra (1 ♂, BMNH) , 47 km W Nyngan at 31°32.8ʹS 146°42.6ʹE (1 ♀, CAS) , 4 km W Sunny Corner at 33°22.7ʹS 149°51.6ʹE (6 ♀, 4 ♂, CAS) , Sydney (1 ♀, 1 ♂, AMS; 1 ♀, BMNH), Sydney: Australian Museum (1 ♀, AMS) , Sydney : Kangaroo Park (1 ♀, UCD) , Sydney : North Ryde (1 ♀, AMS) , 23 km SE Tamworth (2 ♀, 1 ♂, ANIC) , 50 km NW Taree at 31°31ʹS 152°14ʹE (2 ♂, AMS) , Tipaminka (1 ♀, AMS) , Tooloom Plateau via Urbenville (1 ♀, ANIC) , Towra Point Nature Reserve (1 ♀, AMS) , Tuglow River : Mount Werong Fire Road (2 ♀, 1 ♂, AMS) , Warrenburg National Park (1 ♀, UCD) , Warrumbungle National Park at 31°16.9ʹS 148°59.1ʹE (3 ♀, 5 ♂, CAS) , Warrumbungle National Park : Camp Pincham (6 ♀, ANIC) , near Warrumbungle National Park at 31°16.9ʹS 149°04.8ʹE (5 ♂, CAS) , 12.5 km W Wilcannia at 31°39.7ʹS 143°26.0ʹE (1 ♀, CAS) , Wollemi National Park (northern edge) at 32°23.4ʹS 150°24.8ʹE (13 ♀, CAS) . Norfolk Island: Highland Guesthouse at 29°02ʹS 167°57ʹE (2 ♂, ANIC) , Kingston (1 ♀, BISH) , Philip Island near Norfolk Island (2 ♀, CAS) , Rocky Point Reserve at 29°03ʹS 167°55ʹE (1 ♀, 2 ♂, ANIC; 2 ♂, CAS) . Northern Territory : Berry Springs Park 50 km SE Darwin (1 ♂, NTM) , Black Point on Cobourg Peninsula at 11°09ʹS 132°09ʹE (1 ♀, ANIC) , 4 mi W Coolibah Homestead at 15°34ʹS 130°54ʹE (1 ♀, ANIC) , Darwin (1 ♀, NTM) , Gregory National Park at 160°6.6ʹS 130°25.7ʹE (1 ♀, ANIC) , Kakadu National Park (2 ♀, CAS) , Kakadu National Park : Leichardt Gallery in Deaf Adder Valley (1 ♀, ANIC) , Katherine Gorge near Katherine (1 ♀, QMB) , Keep River National Park at 16°03ʹ01ʺS 130°24ʹ07ʺE (1 ♀, CAS) , Mango Plantation at 12°52ʹS 130°35ʹE (1 ♀, NTM) , 19 km NE Mount Cahill at 12°50ʹS 132°52ʹE (1 ♂, ANIC) . Queensland : Arcadia on Magnetic Island at 19°09ʹS146°52ʹE (10 ♀, 3 ♂, ANIC) , Armstrong Beach ca 15 km E Sarina at 21°27.3ʹS 149°17.5ʹE (1 ♀, 2 ♂, CAS) , Atherton at 17°17ʹS 145°29ʹE (5 ♀, 3 ♂, ANIC) , near Atherton (1 ♀, CAS) , 4 km S Ayr at 19°38.2ʹS 147°23.3ʹE (1 ♀, CAS) , 8 km NW Bald Hill in Ilwraith Range at 13°45ʹS 143°22ʹE (1 ♀, ANIC) , Batavia Downs at 12°40ʹS 142°39ʹE (1 ♂, ANIC) , 5 km S Batavia Downs at 12°41ʹS 142°41ʹE (2 ♂, ANIC) , The Bend 3 km NW Coen 13°56ʹS 143°12ʹE (1 ♀, ANIC) , Biggenden (1 ♀, ANIC) , Bin Bin Range (1 ♀, ANIC) , Bluff Range near Biggenden (1 ♀, ANIC) , Bowling Green Bay National Park at 19°26.0ʹS 146°56.7ʹE (14 ♀, 2 ♂, CAS) , Box Creek 16 km N Proserpine (1 ♀, AMS) , Brisbane (2 ♂, RMNH; 10 ♀, 1 ♂, QMB), Brisbane: Bardon (1 ♀, 1 ♂, BMNH) , Brisbane : Blunder Creek (4 ♀, QMB) , Brisbane : Botanic Garden at 27°28.8ʹS 152°58.1ʹE (1 ♀, CAS) , Brisbane : Indooroopilly (2 ♀, 1 ♂, BMNH) , Brisbane : Karawatha Forest at 27°38.6ʹS 153°04.2ʹE (4 ♀, CAS) , Brisbane : Mount Coot-tha (1 ♀, CAS) , Brookfield near Brisbane (8 ♀, BMNH) , Bundaberg (1 ♂, ANIC; 1 ♀, 1 ♂, BMNH), Burdekin River 20 km NE Charters Towers at 20°00.1ʹS 146°26.3ʹE (2 ♂, CAS) , Burnett River at Bundaberg (1 ♀, ANIC) , Cairns (1 ♀, 1 ♂, BMNH) , Caloundra beach at 26°47.1ʹS 153°08.4ʹE (1 ♀, CAS) , Carnarvon National Park at 25°04.0ʹS 148°14.7ʹE (4 ♀, CAS) , Chili Beach near Portland Roads (1 ♀, AMS) , Claudie River near Mount Lammont (1 ♀, AMS) , Coen at 13°57ʹS 143°12ʹE (2 ♀, ANIC) , Conway National Park at 20°17.1ʹS 148°25.8ʹE (3 ♀, CAS) , Cooktown at 15°28.3ʹS 145°15.5ʹE (1 ♂, AMS; 1 ♂, CAS), Crater Lakes National Park via Coalstoun Lakes SW Biggedden (2 ♀, 1 ♂, ANIC) , Crediton State Forest at 21°11.8ʹS 148°29.7ʹE (6 ♀, 2 ♂, CAS) and 21°11.9ʹS 148°29.9ʹE (4 ♀, CAS) , Curtain Fig 2 km SSW Yungaburra at 17°17ʹS 145°34ʹE (1 ♀, ANIC) , Davies Creek National Park at 17°00.2ʹS 145°34.1ʹE (1 ♂, CAS) , 9 km S Dingo Beach at 20°05.5ʹS 148°30.2ʹE (2 ♀, CAS) , 18 km S Dingo Beach at 20°16.0ʹS 148°31.2ʹE (1 ♀, CAS) , Dipperu National Park at 21°53.9ʹS 148°46.5ʹE (5 ♀, CAS) , Edungalba (1 ♀, ANIC) , Emerald (3 ♀, ANIC) , Eungella National Park at 21°10.5ʹS 148°30.3ʹE (61 ♀, 16 ♂, CAS; 7 ♀, 1 ♂, QMB), Eurimbulla National Park at 24º10ʹS 151º52ʹE (1 ♂, AMS) , Fletcher Creek 43 km NW Charters Towers at 19°48.9ʹS 146°03.3ʹE (9 ♀, CAS) , Gwinganna 6 km SSW Tallebudgera at 28°11ʹS 153°23ʹE (1 ♀, ANIC) , Heathlands at 11°45ʹS 142°35ʹE (1 ♀, 3 ♂, ANIC) , 12 km NE Heathlands at 11°43ʹS 142°41ʹE (1 ♀, ANIC) , 12 km SSE Heathlands at 11°51ʹS 142°38ʹE (6 ♀, ANIC) , Homevale National Park at 21°26.9ʹS 148°32.4ʹE (14 ♀, 1 ♂, CAS) , 14 km NW Hope Valley Mission at 15°16ʹS 144°59ʹE (2 ♀, 2 ♂, ANIC) , Horse Gully at foot of Bunya Mountains at 26°42ʹS 150°31ʹE (1 ♂, ANIC) , Iron Range National Park : Middle Cloudie River (4 ♀, AMS) , Isaac River 100 km NE Clermont (1 ♀, QMB) , Kuranda (1 ♀, ANIC; 1 ♀, CAS), Kuranda : Russet Park (4 ♀, 1 ♂, CAS) , 5 km NE Leyburn (2 ♀, CAS) , Mackay (6 ♀, 3 ♂, BMNH) , Mareeba (1 ♀, ANIC) , 65 km N Marlborough (1 ♀, AMS) , Maryborough at 25°32ʹS 152°44ʹE (1 ♀, ANIC) , Mary Creek 14 mi. N Mount Molloy (1 ♀, CAS) , 13 km E Miles at 26°40ʹS 150°19ʹE (1 ♂, ANIC) , Mornish (1 ♀, CAS) , Mossman (1 ♀, CAS) , 56 road km WNW Mount Carbine at 16°19.4ʹS 144°43.2ʹE (1 ♂, CAS) , Mount Lewis 3200 feet (1 ♀, CAS) , 48 km E Mount Surprise at 18°09.0ʹS 144°43.6ʹE (4 ♀, CAS) , Mount Tambourine (1 ♂, BMNH) , Mount Tibrogargane (1 ♀, QMB) , Mount Walsh National Park via Biggenden (1 ♂, ANIC) , 3 km NE Mount Webb at 15°03ʹS 145°09ʹE (1 ♂, ANIC) , Mulgrave River (1 ♀, CAS) , Murrays Spring in Lawn Hill National Park at 18°35ʹ15ʺS 138°04ʹ28ʺE (1 ♀, ANIC) , Musselbrook Camp at 18°36ʹS 138°08ʹE (1 ♀, ANIC) , Normanby River at 15°18ʹS 144°57ʹE (2 ♀, ANIC) , Paluma Range National Park at 18°51.6ʹS 146°07.6ʹE, alt. 50 m (3 ♀, CAS) , Pendland at 20°31.0ʹS 145°24.2ʹE (1 ♀, CAS) , Pinnacle Creek 27 km N Archer Crossing (1 ♀, ANIC) , Port Douglas at 16°31.1ʹS 145°28.3ʹE (10 ♀, 3 ♂, CAS) , Rainforest CRC [= Cooperative Research Centre] at 16°06ʹ16ʺS 145°26ʹ58ʺE (1 ♀, AMNH) , Rockhampton (1 ♀, ANIC) , 2 km N Rokeby at 13°39ʹS 142°40ʹE (4 ♀, 1 ♂, ANIC) , 2 km W Rolleston at 24°27.6ʹS 148°36.2ʹE (1 ♀, CAS) , Shiptons Flat at 15°47ʹS 145°14ʹE (7 ♂, ANIC) , Somerset Dam (2 ♀, QMB) , Split Rock 14 km SE Laura at 15°39ʹS 144°31ʹE (89 ♀, 18 ♂, ANIC; 1 ♂, CAS) and at 15°39ʹS 144°42ʹE (7 ♀, ANIC) , Stanthorpe (1 ♀, QMB) , Tannum Sands at 23°56.8ʹS 151°22.5ʹE (1 ♂, CAS) , Tibrogargan Creek (1 ♂, QMB) , Toowoomba (1 ♂, QMB) , Townsville (1 ♀, BMNH; 1 ♀, RMNH; 4 ♀, 3 ♂, SAM), 11 km S Townsville at 19°21.8ʹS 146°53.2ʹE (5 ♀, 1 ♂, CAS) , 37 km S Townsville at 19°22.4ʹS 147°01.7ʹE (1 ♀, CAS) , Tully (1 ♂, RMNH) , 13 km SE Weipa at 12°40ʹS 143°00ʹE (4 ♀, 2 ♂, ANIC) , Whitsunday Islands (1 ♀, RMNH) , Wonga Beach 11 km NNE Mossman at 16°19.9ʹS 145°25.3ʹE (1 ♀, 2 ♂, CAS) . South Australia : Eckerts Creek at 34°20ʹS 140°34ʹE (1 ♀, ANIC) , Highgate near Adelaide (1 ♀, SAM) , Kangaroo Island : Gosse area (3 ♀, BMNH) , Mitcham near Adelaide (4 ♀, SAM) , 5 km S Mylor (2 ♂, BMNH) , Wilpena in Flinders Ranges National Park at 31°31.7ʹS 138°36.2ʹE (27 ♀, 14 ♂, CAS) , 3 km ENE Wilpena in Flinders Ranges National Park at 31°31.0ʹS 138°36.6ʹE (1 ♀, CAS) , Wilpena Pound Gap at 31°33ʹS 138°36ʹE (1 ♂, ANIC) . Tasmania : Bathurst Harbour at 43°22ʹS 146°08ʹE (1 ♀, ANIC) , 14 km S Bronte Park at 42°15ʹS 146°29ʹE (1 ♀, ANIC) , 12 km NNE Bronte Park at 42°02ʹS 146°33ʹE (1 ♂, ANIC) , 9 km SW Bronte Park at 142°12ʹS 146°30ʹE (2 ♀, ANIC) , Condominium Creek 5 km WSW Mount Anne at 42°58ʹS 146°22ʹE (1 ♂, ANIC) , 5 km ENE Cranbrook at 41°59ʹS 148°07ʹE (1 ♂, ANIC) , 9 km WSW Derwent Bridge at 42°10ʹS 146°08ʹE (1 ♀, ANIC) , 3 km E Dover (1 ♀, CAS) , Edwards Road in Hartz Mountains at 43°07ʹS 146°47ʹE (2 ♀, ANIC) , Ewart Creek at 41°58ʹS 145°28ʹE (2 ♀, ANIC) , Franklin Road at 42°13ʹS 146°01ʹE (3 ♀, ANIC) , 7 km S Frodshams at 42°53ʹS 146°22ʹE (1 ♂, ANIC) , 13 km W Geeveston (1 ♀, BMNH) , 1 km SSE Gladstone (1 ♀, 2 ♂, ANIC) , 5 km SE Harford at 41°15ʹS 146°36ʹE (2 ♂, ANIC) , Hellyer Gorge at 41°16ʹS 145°37ʹE (6 ♀, ANIC) , Hobart (1 ♀, SAM) , Hobart : Sandy Bay (1 ♀, ANIC) , Intake Bridge at 41°19ʹS 147°56ʹE (1 ♀, 1 ♂, ANIC) , Launceston (5 ♀, 2 ♂, SAM) , Launceston : Newstead (2 ♀, ANIC) , The Lea at 42°56ʹS 147°19ʹE (1 ♀, ANIC) , 9 km SE Miena (3 ♀, UCD) , 11 km E Mount Barrow at 41°23ʹS 147°25ʹE (1 ♀, ANIC) , Mount Field National Park (3 ♀, BMNH) , 4 km W Orford at 42°34ʹS 147°50ʹE (1 ♂, ANIC; 1 ♂, CAS), Pelion Gap 2 km ENE Mount Ossa at 41°52ʹS 146°03ʹE (1 ♀, ANIC) , Pelion Hut 3 km S Mount Oakleigh at 41°50ʹS 146°03ʹE (8 ♀, 3 ♂, ANIC) , Pittwater (1 ♀, ANIC) , Poatina at 41°49ʹS 146°54ʹE (4 ♀, ANIC) , 9 km SW Poatina at 41°48ʹS 146°52ʹE (1 ♀, ANIC) , Pyengana (1 ♀, SAM) , 4 km E Rosebery at 41°47ʹS 145°35ʹE (2 ♀, ANIC) , Saint Patricks River (2 ♀, SAM) , 4 km WSW Tim Shea (1 ♂, ANIC) , University of Tasmania Campus at 42°56ʹS 147°21ʹE (3 ♀, ANIC) , 3 km ENE Wayatinah at 42°22ʹS 146°29ʹE (1 ♂, ANIC) , 4 km SE Weldborough at 41°14ʹS 147°56ʹE (1 ♀, 1 ♂, ANIC) , 14 km SSW Wilmot at 41°30ʹS 146°05ʹE (1 ♂, ANIC) , no specific locality (2 ♀, OXUM, lectotype and paralectotype of Pison obliquum ) . Victoria : Balwyn (1 ♀, ANIC) , Crib Point at 38°21ʹ47ʺS 145°12ʹ07ʺE (1 ♀, WAM) , Melbourne : Botanic Garden (1 ♂, BMNH) , 23 mi E Orbost (1 ♀, CAS) , Thomson State Forest 5 km S Aberfeldy (1 ♂, CAS) . Western Australia : Applecross at 32°00ʹ50ʺS 115°50ʹ20ʺE (1 ♀, WAM) , Augusta at 34°18ʹ43ʺS 115°09ʹ32ʺE (1 ♀, WAM) , Busselton (2 ♀, UCD) , Chidlow at 31°51ʹ25ʺS 116°16ʹ19ʺE (1 ♀, WAM) , Denmark (1 ♀, WAM) , Dongarra (1 ♀, BMNH) , Fitzgerald River National Park at 33.949416°S 119.926086°E (10 ♀, 1 ♂, MNKB) , Graylands (1 ♂, WAM) , 12 km S Kalumburu Mission at 14°25ʹS 126°38ʹE (1 ♂, ANIC) , 14 km SE Kalumburu Mission at 14°25ʹS 126°40ʹE (2 ♀, ANIC) , 4 km W King Cascade at 15°38ʹS 125°15ʹE (1 ♂, ANIC) , Lone Dingo on Mitchell Plateau at 14°35ʹS 125°45ʹE (1 ♀, ANIC) , Millstream (1 ♂, ANIC) , Mining Camp on Mitchell Plateau at 14°49ʹS 125°50ʹE (1 ♀, ANIC) , Perth (1 ♀, BMNH) , Perth : Cottesloe (8 ♀, 2 ♂, WAM) , Perth : Darling Range (5 ♀, BMNH) , Perth : Darlington at 31°54ʹ04ʺS 116°04ʹ52ʺE (1 ♀, WAM) , Perth : Maylands (8 ♀, 3 ♂, WAM) , 100 km WSW Point Malcolm at 33°48ʹS 123°46ʹE (2 ♀, 1 ♂, WAM) , Porongorup National Park (1 ♀, CAS) , Serpentine Falls National Park at 32°22.1ʹS 116°00.5ʹE (1 ♀, CAS) , South Dandalup Dam 17 km E North Dandalup at 32°38.7ʹS 116°02.4ʹE (4 ♂, CAS) , South Perth (1 ♀, WAM) , Stokes National Park 66 km W Esperance at 33°49.8ʹS 121°08.4ʹE (1 ♂, CAS) , Yalgorup National Park at 32°54.8ʹS 115°42.1ʹE (1 ♀, CAS) , Yallingup (1 ♂, UCD) , Yanchep 32 mi N Perth (10 ♀, 2 ♂, BMNH) .

COOK ISLANDS: Island of Aitutaki: Aiutaki (1 ♂, BISH) , Amuri (2 ♂, BISH) , no specific locality (3 ♀, BISH) . Island of Rarotonga: Arorangi at 21°12.4ʹS 159°49.4ʹW (1 ♀, 1 ♂, CAS) , Titikaveka (1 ♀, 1 ♂, BISH) , no specific locality (1 ♀, BMNH) .

FEDERATED STATES OF MICRONESIA (Krombein, 1949b or as indicated): Dublon Island : Truk Atoll: Truk and Truk-Erin (Yasumatsu, 1953), Truk: Moen (1 ♀, CAS) . Kosrae Island (formerly Kusaie Island): Lelo and Mwot-Utwe ( Yasumatsu , 1953), Mount Tafeyät , Mutunlik (7 ♀, BISH) . Palau: Peleliu Island. Pohnpei Island (formerly Ponape): Kolonia (5 ♀, BISH) , Metalanum (1 ♀, BISH) , Nanponmal (1 ♀, BISH) , Ronkiti-One (Yasumatsu, 1953), Tamon. Yap Island: no specific locality (1 ♀, BISH) .

FIJI: Cicia: Mabula (1 ♂, BISH) . Ovalau: Levuka (1 ♀, 1 ♂, BISH) . Vanua Balavu: Nabavatu (1 ♂, BISH) . Vanua Levu: Savusavu Estate (4 ♂, CAS) . Viti Levu: Korotogo 8 km E Sigatoka (1 ♀, CAS) , Nadi (1 ♀, BISH) , 10 km E Sigatoka (1 ♀, 4 ♂, CAS) , Suva (2 ♀, CAS; 10 ♀, 3 ♂, RMNH) .

FRENCH POLYNESIA (Cheesman, 1928; Williams, 1932, or as indicated): Moorea: Atitia (4 ♂, CAS) , Baie de Cook (2 ♀, BISH) . Marquesas Islands: Fatu-hiva; Hiva-oa: Atuona Valley; Nuku-hiva: Teivipakeka; Tahuata: Kiinui Valley; Ua-huka: Hane Valley ; Ua-pou: Hakahetau . Society Islands: Raiatea: Uturoa (2 ♀, BISH) , Vaitape (1 ♀, 1 ♂, BISH) . Tahiti: between Lake Vaihiria and Otiaroa road (Menke, 1979), Tairapu . Tubuai Island: Mahu (2 ♀, BISH) .

HAWAIIAN ISLANDS: Hawaii: Kapua Bay (1 ♀, BISH) . Maui: Haiku (9 ♀, BISH) , Makena (3 ♀, BISH) . Molokai (Krauss, 1944): Molokai , south central Molokai. Niihau : no specific locality (1 ♀, BISH) . Oahu: Halona Valley at 21°25ʹN 158°06ʹW (1 ♂, CAS) , Honolulu: Ewa (2 ♂, BISH; 1 ♀, 1 ♂, CAS), Honolulu (2 ♀, BISH; 1 ♂, BMNH, lectotype of Pison iridipenne ; 1 ♀, UCD), Honolulu: Nuuanu Valley (1 ♀, CAS) , Honolulu: Punchbowl (1 ♂, CAS) , Honolulu: Woodlawn (1 ♀, 2 ♂, CAS) , Manoa (1 ♀, BISH) , Mount Tantalus (1 ♀, BISH) , Waipio (1 ♂, BISH) .

INDONESIA: Ambon: no specific locality (5 ♀, RMHN) . Halmahera: between Payahe and Gita Woda (2 ♂, RMNH) , Sagutora 20 km S Payahe (1 ♂, RMNH) . Java: Bogor (1 ♀, RMHN) , Jakarta (2 ♀, 1 ♂, as Batavia, RMNH) . Seram: 11 km E Wahai (1 ♀, RMHN) . Sula: island of Mangoli: near Buya (1 ♂, RMNH) , island of Taliabu: near Semuya (1 ♂, RMNH) . Sulawesi: Mount Bulusaraung: near Camba Malawa (1 ♂, RMNH) . Sumatra: western Sumatra : no specific locality . Western Papua (= Indonesian New Guinea): Island of Biak: Bosnik at 1°09ʹS 136°14ʹE (1 ♂, RMNH) , Kampong Landbouw (1 ♀, BISH) , and Sorido (1 ♀, AMNH) , Ifar (1 ♀, RMHN) , Island of Yapen: Davai River SSE Sumberbaba (1 ♀, 1 ♂, BISH, as Japen), Jayapura (14 ♀, 3 ♂, RMHN, as Hollandia), Karubaka in Swart Valley at 3°36ʹS 138°28ʹE (2 ♀, BISH) , Kebar Valley W Manokwari (1 ♀, BISH) , Maffin Bay (1 ♀, 2 ♂, CAS) , Mimika River (1 ♀, BMNH, holotype of Pison impunctatum ), Mount Gyifrie (1 ♀, SAM) , Paniai Lakes (as Wisselmeren): Enarotadi (2 ♀, BISH) , Moanemani (1 ♀, BISH) and Urapura (1 ♀, BISH) , Sentani at 2º40ʹS 140º30ʹE (2 ♂, RMNH) , Sibil Valley in Star Mountains at 5º00ʹS 141º00ʹE (1 ♀, BISH) , mouth of Tor River 4 km E Hol Maffen (1 ♀, 2 ♂, BISH) .

JAPAN: Ogasawara (= Bonin) Islands: Chichijima Island: Sakai-ura (1 ♂, BISH), Hahajima Island (1 ♀, BISH) .

KIRIBATI REPUBLIC: Teraina Island (1 ♂, BISH, as Washington Island ) .

MALAYSIA: Sabah: near Penampang (2 ♂, CAS) , Ranau (2 ♀, BISH) , Singkor (1 ♂, BISH) . Sarawak: Meirai in Kapit District (1 ♀, BISH) . Selangor: Dusun Tua in Hulu Langat area (1 ♀, RMNH, as Ulu Langat) .

MARIANA ISLANDS (Krombein, 1949b or as indicated): Guam: Ritidian Point (2 ♂, BISH), no specific locality (1 ♂, BISH). Saipan: Saipan Island (5 ♂, BISH), Tanapag (Krombein, 1950). Tinian Island: Hagoi Lake, Marpo Valley, Mount Lasso .

MARSHALL ISLANDS: Ailinglaplap Atoll: Bigatyelang Island (12 ♀, BISH) , Arno Atoll: Ine Island (3 ♀, 2 ♂, BISH) , Ebon Island (3 ♀, BISH) , Jaluit Atoll ( Yasumatsu , 1953), Wotho Atoll (25 ♀, 2 ♂, BISH) , Wotje Atoll (1 ♂, BISH) .

NEW CALEDONIA: Grande Terre: Anse Vata (1 ♂, BISH) , Mont Koghi (1 ♀, BISH) , Nouméa (2 ♀, ANIC; 7 ♀, 6 ♂, BISH), Oua Tom (1 ♂, BISH) , Saint Louis, a part of Mont Doré village (2 ♂, BISH) , Thio River valley (1 ♀, BISH) , Touho-Houaïlou (1 ♀, BISH) .

PALAU REPUBLIC: Koror Island (13 ♀, 7 ♂, BISH; 1 ♀, ELKU, holotype of Pison korrorensis ; 1 ♂, USNM) .

PAPUA NEW GUINEA: Central Province: Bome in Goilala District (2 ♀, BISH) , Rouna Falls 46 km E Port Moresby (1 ♀, BISH) . Eastern Highlands Province: Aiyura (1 ♂, BISH) , Kainantu (1 ♀, BISH) , Kassam 48 km E Kainantu (1 ♀, 1 ♂, BISH) , 22 km SE Okapa (1 ♀, BISH) . East Sepik Province: Angoram (1 ♀, BISH) . Island of Bougainville: Buin (1 ♀, BISH) , Buka (1 ♀, BISH) . Madang Province: Aranam 2 air km W Bundi at 5°45ʹS 145°15ʹE (3 ♀, 12 ♂, CAS) , Baiteta 12 km NW Alexishafen at 5°00ʹS 145°45ʹE (3 ♀, 1 ♂, CAS) , Batua 2 air km SW Bundi at 5°45ʹS 145°15ʹE (4 ♀, CAS) , Brahman Catholic Mission at 6°45ʹS 145°23ʹE (1 ♂, CAS) , Bundi at 5°45ʹS 145°15ʹE (36 ♀, 41 ♂, CAS) , Erima (1 ♀, MTM, determined P. iridipenne by Tsuneki), Gogol River 12 km SW Madang at 5°20ʹS 145°42ʹE (1 ♀, CAS) , Karisokora 7.5 km W Bundi at 5°44ʹS 145°10ʹE (1 ♀, CAS) , Karkar Island: Bagiai Crater Trail (1 ♀, BISH) and Kevasop village (2 ♀, CAS) , 5 air km NE Mundiai Pass at 5°46ʹS 145°09ʹE, alt. 2,500 m (1 ♀, CAS) , Nagada Harbor 8 km N Madang at 5°09ʹS 145°48ʹE (2 ♀, 10 ♂, CAS) , Nobonob Hill 7 km NW Madang at 5°10ʹS 145°45ʹS (11 ♀, 14 ♂, CAS) , Pandambai 6 air km W Bundi at 5°38ʹS 145°11ʹE, 2,330 m. alt. (5 ♀, CAS) , Saidor: Kiambavi (1 ♀, BISH) , Sapi Forest Reserve 30 km W Madang at 5°12ʹS 145°30ʹE (1 ♂, AMNH; 25 ♀, 58 ♂, CAS), Simbai at 5°17ʹS 144°26ʹE (1 ♀, AMNH; 16 ♀, 31 ♂, CAS), Snow Pass at 5°44ʹS 145°18ʹE (1 ♀, CAS) , Tapo Creek 26 km SW Madang at 5°24ʹS 145°38ʹE (3 ♀, 2 ♂, CAS) Wanuma (1 ♀, BISH) . Milne Bay Province: Normanby Island: Wakaiuna (1 ♀, BISH) , Woodlark Island: Kulumadau Hill (1 ♀, BISH) . Morobe Province: Busu River near Lae (1 ♀, BISH) , Finschhafen (2 ♀, BISH) , Mindik (1 ♂, BISH) , Mount Kaindi, 2,440 m (1 ♂, BISH; 1 ♀, CAS), Mount Missim at 7º15ʹS 146º48ʹE (1 ♀, 1 ♂, BISH) , Pindiu (1 ♀, BISH) , Wau (44 ♀, 22 ♂, BISH; 2 ♀, 1 ♂, RMNH), Wau: Edie Creek (1 ♀, BISH) , Wau: Kunai Creek (1 ♀, BISH) .

National Capital District: Port Moresby (1 ♂, BISH; 3 ♀, 1 ♂, CAS) . New Britain (Tsuneki, 1982, or as indicated): Hermit Islands: Luf ; Keravat (1 ♀, BISH) , Valoka, Yalom. New Ireland: Hans Meyer Range (1 ♀, AMS) , Lemkamin (Tsuneki, 1982), Kavieng (1 ♀, 1 ♂, BISH) . Oro Province: Kokoda (1 ♀, BMNH) , Popondetta (1 ♀, BISH) . Sandaun Province (= West Sepik): island of Seleo (1 ♂, MTM, determined P. iridipenne by Tsuneki), Torricelli Mountains (1 ♀, UCD) . Southern Highlands Province: Dimifa (1 ♀, BISH) , Koroba 40 km W Tari (2 ♀, BISH) . Western Highlands Province: Nondugl (1 ♀, BISH) .

PHILIPPINES: Luzon: Buguias 60 km S Bontoc (2 ♀, BISH) , Pili on Mount Isarog (1 ♂, BISH) , Wawa Dam (1 ♀, 1 ♂, BISH) . Mindanao: Jolo Island : 8-10 km S Taglibi (1 ♀, BISH) , Mount Pamalihi 21 km W Gingoog (2 ♀, 1 ♂, BISH) . Palawan: Pinigisan (Tsuneki, 1976) .

PITCAIRN ISLANDS: Pitcairn Island (1 ♀, CAS) .

SAMOA: Upolu: Mount Vaea (1 ♀, BISH), no specific locality (1 ♀, RMNH) ,

SINGAPORE: Singapore (1 ♀, 2 ♂, CAS) .

SOLOMON ISLANDS: Bellona: western end (1 ♀, BISH) . Guadalcanal: Paripao (1 ♀, BISH) . Kolombangara: Pepele (1 ♀, BISH) . Malaita: Tagatalau (1 ♀, BISH) . New Georgia: Munda (1 ♂, BISH) . Rennel: Te-Uhunhango (1 ♂, BISH) . Santa Cruz: Graciosa Bay (1 ♀, BISH) . Tulagi: Chinatown (4 ♀, RMNH) , locality illegible (1 ♀, RMNH) , no specific locality (1 ♀, 1 ♂, RMNH) .

THAILAND: Ton Nga Chang National Park 27 km SW Hat Yai at 6°54ʹN 100º21ʹE (2 ♀, RMNH) .

TOKELAU ISLANDS: (Hinckley, 1969, as P. iridipenne ).

TONGA: Tongatapu: Nuku’alofa (1 ♀, BISH) , locality unknown (1 ♀, BISH) . Vavau: Neiafu (1 ♀, BISH) .

VANUATU: Efate: Port Vila (2 ♀, 1 ♂, BISH) . Malekula: Lakotoro (2 ♀, BISH) . Tanna: Lenakel (1 ♀, BISH) .

Figures

FIGURES 1-3. Pison abductor Pulawski, sp. nov., female. (1) Clypeus and mandibles; (2) Head in dorsal view; (3) Mandible in lateral view (arrow shows abductor ridge). FIGURE 4. Collecting locality of Pison abductor Pulawski, sp. nov.