Pison rufipes Shuckard

Pulawski, Wojciech J., 2018, A Revision of the Wasp Genus Pison Jurine, 1808 of Australia and New Zealand, New Guinea, and the Pacific Islands (Hymenoptera: Crabronidae), Proceedings of the California Academy of Sciences 65, pp. 1-584 : 396-401

Pison rufipes Shuckard

Figures 960 -966.

Pison rufipes Shuckard, 1838:79 , ♀. Lectotype: ♀, Australia: Van Diemen’s Land, now Tasmania: no specific locality (BMNH), present designation, examined. – F. Smith, 1956:317 (in catalog of Hymenoptera in British Museum); Kohl, 1885:188 (in checklist of world Pison ); Froggatt, 1892:217 (in catalog of Australian Hymenoptera ); Dalla Torre, 1897:713 (in catalog of world Hymenoptera ); Turner, 1915:557 (in key to Pison of Tasmania), 558 (Tasmania), 1916b:596 (in key to Australian Pison ), 602 (recognition characters; Australia: Tasmania); Williams, 1945:438 ( New Caledonia); R. Bohart and Menke, 1976:336 (in checklist of world Sphecidae ); Evans, Matthews, and Hook, 1981:221 (nest and prey); Cardale, 1985:262 (in catalog of Australian Sphecidae ); Callan, 1990:22 ( New Caledonia: no specific locality); K. Walker, Naumann, Austin, Taylor, and Cardale, 1992:49 (in catalog of insects of Tasmania); Naumann, 1993:185 ( Australia: Queensland: Heathlands area in Cape York); Baker, 1998:173 (type origin and depository); Naumann, 1998:185 ( Australia: Queensland: Musselbrook area, approximately 18°40ʹS 138°23ʹE); Jennings, Krogmann, and Burwell, 2013:32 (in checklist of Hymenoptera of New Caledonia). – As Pisonitus rufipes, F. Smith, 1869:298 (new combination, in checklist of Pisonitus ).

LECTOTYPE DESIGNATION AND SPECIES IDENTITY.– Shuckard did not give the number of the specimens examined in his original description of Pison rufipes , but he saw at least two, as indicated by the statement “in the collection of Mr. Westwood and my own”. I have designated as the lectotype the only specimen present in the Natural History Museum, London.

The specimen is in poor condition; in particular, the head, detached from the body and glued on a separate piece of cardboard, is that of a Trypoxylon , as evidenced by the emarginate eye margins in combination with a V-shaped carina above the antennal sockets and a short longitudinal carina emerging from the center of the latter. The remaining body, although partly matted by moisture, has clearly the characteristics of either P. prostratum or P. rufipes (the legs, in particular, are ferruginous). The absence of the head precludes the species recognition, but Shuckard’s original description is of help. He says, in his Latin diagnosis: “ Niger ; mandibulis basi, palpis pedibusque rufis”, and repeats it in the English description: “Black … The mandibles and palpi rufescent”. Since the flagellum is not listed among the rufescent body parts, it must have been black, a character of the species here treated as rufipes .

RECOGNITION.– Like Pison argentatum and P. prostratum , P. rufipes has the head, thorax, propodeum, and gaster all black, the second recurrent vein received near the middle of the second submarginal cell, and integument in most specimens narrowly depressed between postspiracular carina and episternal sulcus. Also, the distance between the antennal socket and orbit is slightly less than socket width, the scutal flange is slightly projecting beyond the anterior margin of the axilla, the posterior scutal margin is slightly concave next to the apex of the flange, and the propodeal dorsum is ridged.

Pison rufipes closely resembles P. argentatum , but differs in having the setae of the upper frons either erect, sinuous ( Fig. 962 ) or suberect, bent ventrally and about as long as 1.0-1.5 × midocellar diameter (in addition to the appressed setae). In P. argentatum , the erect setae of the upper frons are about as long as 0.5 × midocellar diameter. In the female of P. rufipes , the ocellocular distance is 1.2-1.5 × the hindocellar diameter (rather than 0.8-1.1 ×), the clypeal lamella is truncate or broadly obtusely angular, but exceptionally it has a minute median tooth, as in P. argentatum ; the legs are mostly ferruginous, but exceptionally all black, as in P. argentatum . In the male, sternum VIII is punctate and setose well before apex ( Fig. 963 ), whereas in P. argentatum it is unsculptured and asetose except near the hindmargin ( Fig. 82 ).

Unlike P. prostratum , the ocellocular distance of P. rufipes is greater than the distance between the hindocelli or equidistant, the setae of the upper frons and the interocellar area are erect or inclined ventrad (about as long as 1.0-1.5 × midocellar diameter), and the free margin of the clypeal lamella is truncate or broadly angulate in the female. In P. prostratum , the ocellocular distance in the vast majority of specimens is smaller than the distance between hindocelli, the setae are appressed on the upper frons and the interocellar area (in some specimens there are sparse erect setae up to 0.5 × midocellar diameter long), and the free margin of the clypeal lamella is arcuate in most females.

DESCRIPTION.– Frons dull, minutely punctate, punctures shallow, contiguous. Distance between antennal socket and orbit slightly less than socket width. Labrum shallowly, broadly emarginate. Anteromedian pronotal pit transversely elongate, about as long as midocellar diameter. Scutum at most indistinctly foveate along flange, with or without short longitudinal ridges adjacent to posterior margin; scutal and mesopleural punctures shallow, averaging less than one diameter apart; interspaces dull on scutum, shiny on mesopleuron ventrally; scutal flange slightly projecting beyond anterior margin of axilla, posterior scutal margin slightly concave next to apex of flange. Postspiracular carina present, about twice as long as midocellar diameter; integument in most specimens narrowly depressed between postspiracular carina and episternal sulcus; in some specimens, however, the depression is ill defined. Metapleural sulcus costulate between dorsal and ventral metapleural pits. Propodeum with irregular longitudinal carina separating side from dorsum and posterior surface and extending from gastral socket area toward spiracle; dorsum with short oblique ridges emerging from middle carina, otherwise obliquely ridged (ridges becoming larger next to lateral longitudinal carina); side ridged, punctate between ridges; posterior surface ridged. Second recurrent vein ending near middle of submarginal cell II. Posteroventral forefemoral surface microscopically, closely punctate. Hindcoxal dorsum with outer margin not carinate. Punctures of tergum I minute, less than one diameter apart. Sternum II punctate throughout, most punctures slightly more than one diameter apart.

Setae silvery, totally concealing integument on clypeus, appressed on thorax, forecoxal venter, femoral venters, and tergum I; either erect, sinuous or suberect, bent ventrad on upper frons (setal length 1.0-1.5 × midocellar diameter), also with patch of dense, appressed setae below midocellus; suberect on lower gena (setal length about 0.7 × midocellar diameter).

Head, thorax, propodeum, and gaster black, female clypeus ferruginous next to lobe free margin; mandible black basally, yellowish brown subbasally, ferruginous subapically, dark apically; antenna all black or (some males) flagellum yellowish brown ventrally. Femora, tibiae, and tarsi ferruginous in most specimens, but femora and tibiae black in some (e.g., in a female from Mount Kosciuszko National Park at 9,000 feet, 18 females and one male from Canberra and six females from Black Mountain , Australian Capital Territory, a female and a male from Orange Botanic Gardens and three females from 4 km W Sunny Corner, New South Wales, and a male from Orbost , Victoria) .

♀.– Upper interocular distance equal to 0.92-1.04 × lower interocular distance; ocellocular distance equal to 1.2-1.5 × hindocellar diameter, distance between hindocelli 0.9-1.1 × hindocellar diameter; eye height equal to 1.02-1.04 × distance between eye notches. Free margin of clypeal lamella truncate ( Fig. 960 ). Dorsal length of flagellomere I 2.4-2.7 × apical width, of flagellomere IX 1.1-1.2 × apical width. Mandible: trimmal carina with minuscule incision at about one third of length. Length 4.9-7.6 mm; head width 1.7-2.4 mm.

♂.– Upper interocular distance equal to 1.12-1.18 × lower interocular distance; ocellocular distance equal to 1.2-1.4 × hindocellar diameter, distance between hindocelli 1.1-1.2 × hindocellar diameter; eye height equal to 1.00-1.04 × distance between eye notches. Free margin of clypeal lamella obtusely pointed in most specimens ( Fig. 961 ), but arcuate with a round median point in some, or roundly pointed mesally and concave on each side of midpoint (as in P. virosum , see

Fig. 1175). Dorsal length of flagellomere I 1.9-2.0 × apical width, of flagellomere X 0.9-1.0 × apical width; venter of flagellomere III concave basally, convex preapically (inconspicuously so in small specimens). Sternum VIII shallowly, broadly emarginate apically ( Fig. 963 ). Genitalia: Figs. 964, 965 . Length 4.8-6.2 mm; head width 1.6-1.9 mm.

NESTING HABITS.– Evans, Matthews, and Hook (1981) found that this species had two generations near Brisbane, Queensland. They examined three nests from that area. The first two were found on roots dangling from the top of steep, overhanging earthen banks not far from water, and the third on the pendant branch of an Acacia tree about 1.3 m above the ground and close to water. All three were made of dried mud and were spindle-shaped, with the long axis perpendicular to the ground. They were so covered with mud on the outside that individual cells could not be discerned on the surface. The nest on a living branch differed from the other two by having the leaf bases incorporated into its body. The cells measured from 4.0 to 7.5 cm in length and about 2.0- 2.5 cm in width. The third nest was dissected and found to contain 10 cells, all broadly elliptical, 6.5-8.0 mm in diameter and 8.5-12.0 mm in length. Each was separated by at least 2 mm of mud and from the outside of the nest by about 7 mm of mud. The number of spider prey varied from 4 fairly large to 9 small ones per cell. The wasp egg was laid longitudinally, laterally on the base of the opistosoma. The spiders were all Salticidae of the following species: Euryattus bleekeri (Doleschall) , Jotus braccatus L.Koch , and Saitis nigriceps (Keyserling) , now Maratus nigriceps (Keyserling) . GEOGRAPHIC DISTRIBUTION (Fig. 966).–

All Australia including Tasmania, listed from

New Caledonia by Williams (1945), Callan

(1990), and by Jennings, Krogmann, and Burwell (2013).

RECORDS.– AUSTRALIA: Australian Capital

Territory: Black Mountain at 35°16ʹS 149°06ʹE

( 11 ♀, 1 ♂, ANIC; 1 ♂, BMNH; 1 ♀, CAS; 1 ♀,

UCD), Canberra (38 ♀, 5 ♂, ANIC) , Cotter River at

Bendora Creek (1 ♀, CAS). New South Wales:

Armidale (1 ♂, QMB), Burrendong Botanic Garden at 32°42.1ʹS 149°06.2ʹE (1 ♀, CAS), Cabramatta

(1 ♂, BMNH), Cairncross State Forest 15 km

N Wauchope (1 ♀, ASM), Congo 8 km SSE Moruya FIGURE 966. Collecting localities of Pison rufipes Shuckard .

at 35°58ʹS 150°09ʹE (2 ♀, 1 ♂, ANIC), Coolbaggie

Forest Reserve 10 km E Eumungerie at 31°58.5ʹS 148°40.5ʹE (5 ♀, 3 ♂, CAS), 1 km W Eumungerie at 31°56.7ʹS 148°36.9ʹE (1 ♀, CAS), Goonoo State Forest 5 mi. S Mendooran (1 ♀, 1 ♂, AMS), Lorien Wildlife Refuge 3 km N Lansdowne near Taree (1 ♂, AMS), Mudgee (1 ♀, AMS), 40.5 km SW Narrabri at 30°37.7ʹS 149°34.1ʹE (2 ♀, CAS), North Richmond (1 ♂, ANIC), Orange Botanic Garden at 33°15.3ʹS 149°05.7ʹE (1 ♀, 1 ♂, CAS), Pipers Creek in Kosciuszko National Park at 9,000 feet (1 ♀, CAS), 4 km W Sunny Corner at 33°22.7ʹS 149°51.6ʹE (1 ♀, CAS), Sydney (1 ♀, AMS), 23 km SE Tamworth (2 ♂, ANIC), Thirlmere Lakes National Park (1 ♀, UCD), Wahroonga (5 ♀, 3 ♂, AMS), Warrumbungle National Park at 31°16.9ʹS 148°59.1ʹE (1 ♂, CAS) and at 31°16ʹS 148°57ʹE (1 ♀, 2 ♂, MNKB), near Warrumbungle National Park at 31°16.9ʹS 149°04.8ʹE (1 ♀, CAS), Wollemi National Park (northern edge) at 32°23.4ʹS 150°24.8ʹE (1 ♀, CAS). Northern Territory: Buchanan Highway 31 km SSE Victoria Highway at 15°57ʹ37ʺS 130°38ʹ20ʺE (1 ♂, ANIC; 2 ♀, USU) , 14 km NW Cape Crawford at 16°34ʹS 135°41ʹE (1 ♂, ANIC), Charles Darwin National Park in Darwin (1 ♂, NTM), Darwin (1 ♂, NTM), Gregory National Park at 15°36ʹ43ʺS 130°24ʹ08ʺE (5 ♀, ANIC; 1 ♀, CAS; 4 ♀, 1 ♂, USU) , at 15°44ʹ54ʺS 129°10ʹ19ʺE (1 ♂, CAS) , at 15°45ʹ30ʺS 129°06ʹ28ʺE (1 ♀, ANIC; 1 ♂, USU) , at 15°57ʹ33ʺS 129°01ʹ44ʺE (1 ♀, 1 ♂, CAS) , at 15°57ʹ37ʺS 130°38ʹ20ʺE (1 ♂, ANIC) , at 15°57ʹ55ʺS 129°01ʹ52ʺE (1 ♂, ANIC) , at 16°02.4ʹS 130°27.3ʹE (1 ♀, USU), at 16°03.7ʹS 130°27.1ʹE (1 ♀, 1 ♂, ANIC; 1 ♂, CAS; 1 ♂, USU), at 16°06.6ʹS 130°25.7ʹE (1 ♀, 1 ♂, ANIC; 1 ♀, 2 ♂, CAS; 1 ♀, 3 ♂, USU), at 16°06.7ʹS 130°25.4ʹE (2 ♀, 1 ♂, ANIC; 1 ♀, CAS; 4 ♀, USU), at 16°06ʹ42ʺS

130°25ʹ23ʺE (1 ♀, USU) , at 16°07ʹ55ʺS 130°26ʹ11ʺE (2 ♀, ANIC) , at 16°08.9ʹS 130°26.6ʹE (1 ♀, 2 ♂, ANIC; 2 ♂, CAS; 4 ♀, USU), at 16°09.8ʹS 130°26.5ʹE (2 ♀, ANIC) , at 16°10ʹ49ʺS 130°25ʹ51ʺE (4 ♂, ANIC; 3 ♀, USU), and at 16°12ʹ47ʺS 130°25ʹ11ʺE (3 ♀, ANIC; 2 ♂, CAS; 1 ♀, 1 ♂, USU), 12 km S Kalkarindji at 17°31.2ʹS 130°53.8ʹE (2 ♀, 1 ♂, ANIC) , 91 km SW Kalkarindji on Buntine Highway at 17°40ʹ36ʺS 130°00ʹ24ʺE (4 ♀, 4 ♂, USU) , Keep River National Park at 15°44ʹ17ʺS 129°06ʹ55ʺE (3 ♀, 6 ♂, ANIC) , 15°45ʹ30ʺS 129°06ʹ28ʺE (1 ♂, USU) , 15°57ʹ33ʺS 129°01ʹ44ʺE (1 ♂, USU) , 15°57ʹ55ʺS 129°01ʹ52ʺE (1 ♀, ANIC) , 16°03ʹ01ʺS 130°24ʹ07ʺE (1 ♀, ANIC; 2 ♀, USU), Victoria Highway 38.5 km SW Timber Creek at 15°42ʹ40ʺS 130°07ʹ48ʺE (2 ♀, ANIC; 1 ♀, CAS; 1 ♀, USU), Victoria Highway 110 km WSW Timber Creek at 15°56ʹ11ʺS 129°35ʹ22ʺE (3 ♀, 6 ♂, USU) , Victoria Highway at 16°03ʹ22ʺS 129°05ʹ15ʺE (3 ♂, ANIC) , Virginia 31 km SE Darwin Central Business District at 12°33ʹS 131°02ʹE (4 ♀, 1 ♂, NTM) , West MacDonnell National Park: Ellery Creek Big Hole 92 km W Alice Springs at 23°46.7ʹS 133°04.4ʹE (1 ♀, CAS) . Queensland: Bald Mountain area via Emu Vale (1 ♀, QMB) , Balgal Beach 51 km NW Townsville at 19°02.5ʹS 146°25.2ʹE (1 ♂, CAS) , Brisbane (1 ♀, 1 ♂, QMB) , Brisbane: Blunder Creek (10 ♀, 2 ♂, QMB) , Brisbane: Indooroopilly (3 ♀, 1 ♂, BMNH) , Brisbane: Karawatha Forest at 27°38.6ʹS 153°04.2ʹE (1 ♀, CAS) , Brisbane: Mount Coot-tha (1 ♀, CAS) , Bundaberg at Burnett River (1 ♂, ANIC) , Bunya Mountains National Park: Horse Gully Creek (1 ♂, AMS) , Carnarvon National Park at 25°04.0ʹS 148°14.7ʹE (1 ♀, CAS; 1 ♀, QMB), Cockatoo Creek at 11°39ʹS 142°27ʹE (1 ♀, ANIC) , Coen at 13°57ʹS 143°12ʹE (1 ♀, 1 ♂, ANIC) , Crediton State Forest at 21°11.8ʹS 148°29.9ʹE (1 ♀, 1 ♂, CAS) , Dulhunty River 13 km SW Heathlands Homestead at 11°50ʹS 142°41ʹE (1 ♂, QMB) , Esk (1 ♀, QMB) , Eungella National Park at 21°10.5ʹS 148°30.3ʹE (3 ♀, CAS) , Fletcher Creek 43 km NW Charters Towers at 19°48.9ʹS 146°03.3ʹE (1 ♀, CAS) , Goomeri – Petrie Highway 17 (1 ♀, UCD) , Gunshot Creek 13 km NW Heathlands Homestead at 11°43ʹS 141°28ʹE (1 ♀, QMB) , Hann River at 15°11ʹS 143°52ʹE (2 ♀, ANIC) , Heathlands at 11°45ʹS 142°35ʹE (2 ♀, ANIC) , 12 km NE Heathlands at 11°43ʹS 142°41ʹE (2 ♀, ANIC) , Homevale National Park at 21°26.9ʹS 148°32.4ʹE (1 ♀, 3 ♂, CAS) , Kuranda (1 ♀, BMNH) , Kuranda : Russett Park (2 ♀, CAS) , Lake Monduran at 24°52.1ʹS 151°51.0ʹE (1 ♀, CAS) , Lawn Hill (now Boodjamulla ) National Park at 18°35ʹ15ʺS 138°04ʹ28ʺE (1 ♀, QMB) and 18°40ʹ15ʺS 138°22ʹ15ʺE (6 ♀, QMB) , Maryborough at 25°32ʹS 152°44ʹE (1 ♀, ANIC) , 48 km E Mount Surprise at 18°09.0ʹS 144°43.6ʹE (3 ♀, CAS) , Mount Tibrogargan (3 ♀, 2 ♂, QMB) , Murrays Spring 8 km NW Musselbrook Camp at 18°35ʹS 138°03ʹE (5 ♀, ANIC) , Musselbrook Camp at 18°36ʹS 138°08ʹE (1 ♂, ANIC) , Rocky Creek 44 km N Moreton in York Peninsula (1 ♀, ANIC) , 2 km N Rokeby at 13°39ʹS 142°40ʹE (1 ♀, 1 ♂, ANIC) , 61 km S Rolleston at 24°59.7ʹS 148°27.8ʹE (3 ♀, 2 ♂, CAS) , Somerset Dam (4 ♀, QMB) , 6 km N Taroom at 25°36ʹS 149°46ʹE (1 ♀, QMB) , Townsville (1 ♀, RMNH) , 13 km SE Weipa at 12°40ʹS 143°00ʹE (2 ♀, ANIC) . South Australia: Adelaide (1 ♂, SAM) , Wilpena in Flinders Ranges National Park at 31°31.7ʹS 138°36.2ʹE (25 ♀, 12 ♂, CAS) , 3 km ENE Wilpena at 31°31.0ʹS 138°36.6ʹE (22 ♀, 3 ♂, CAS) , Wilpena Pound Gap at 31°35ʹS 138°36ʹE (1 ♀, ANIC) . Tasmania: 3.1 km N Bronte Park (1 ♀, CAS) , 12 km NNE Bronte Park at 42°02ʹS 146°33ʹE (3 ♀, ANIC) , 14 km S Bronte Park at 42°15ʹS 146°29ʹE (1 ♀, 1 ♂, ANIC) , Collinsville (1 ♀, BMNH) , 9 km WSW Derwent Bridge at 42°10ʹS 146°08ʹE (1 ♀, ANIC) , Edwards Road in Hartz Mountains at 43°07ʹS 146°47ʹE (1 ♀, ANIC) , Geeveston Park (1 ♀, BMNH) , 1 km SSE Gladstone at 40°58ʹS 148°01ʹE (1 ♀, ANIC) , Great Pine Tier 13 km NNW Bronte Park (3 ♀, BMNH) , Hobart (1 ♀, SAM) , Launceston (1 ♀, ANIC) , 9 km SE Miena (3 ♀, UCD) , Mount Field National Park (1 ♀, ANIC; 2 ♀, BMNH), Pelion Hut 3 km S Mount Oakleigh at 41°50ʹS 146°03ʹE (1 ♀, ANIC) , Poatina at 41°49ʹS 146°54ʹE (8 ♀, ANIC) , 3 km ENE Wayatinah at 42°22ʹS 146°29ʹE (1 ♂, ANIC) , no specific locality (1 ♀, BMNH, lectotype of Pison rufipes ) . Victoria: Melbourne (1 ♀, BMNH) , 18 km NNW Omeo (1 ♀, ANIC) , 23 mi. E Orbost (1 ♂, CAS) . Western Australia: Boya (1 ♀, WAM) , Carson escarpment at 14°49ʹS 126°49ʹE (1 ♂, ANIC) , 10 km W Cobra Station at 24°10.2ʹS 116°23.0ʹE (23 ♀, 14 ♂, ANIC; 2 ♀, USU), 22 km E Cobra Station at 23°13.3ʹS 116°33.1ʹE (14 ♀, 7 ♂, USU) , Fitzgerald River National Park at 33.949416°S 119.926086°E (1 ♂, MNKB) , Great Northern Highway at 23°07.3ʹS 119°05.5ʹE (1 ♀, ANIC) , Karijini National Park at 22°25.6ʹS 118°23.7ʹE (1 ♂, USU) , at 22°26.3ʹS 118°22.9ʹE (1 ♀, ANIC) , at 22°28.4ʹS 118°32.6ʹE (1 ♀, ANIC; 1 ♀, USU), and at 22°28.8ʹS 118°21.6ʹE (3 ♀, 1 ♂, ANIC) , Kennedy Range National Park at 24°38.7ʹS 115°10.7ʹE (1 ♀, ANIC; 1 ♀, CAS; 1 ♀, USU), 28 km E Leonora (1 ♀, CAS) , 11 km E Marble Bar at 21°09.0ʹS 119°51.7ʹE (1 ♂, ANIC) , 30 km E Marble Bar at 21°11.0ʹS 120°01.7ʹE (2 ♀, ANIC; 1 ♂, USU), 63 km E Marble Bar at 21°13.0ʹS 120°20.2ʹE (3 ♀, ANIC; 1 ♂, USU), 95 km E Marble Bar at 21°16.8ʹS 120°36.3ʹE (1 ♀, USU) , 104 km E Marble Bar at 21°19.1ʹS 120°40.3ʹE (4 ♀, ANIC) , 133 km SW Marble Bar at 21°41.6ʹS 119°04.8ʹE (16 ♀, 3 ♂, USU) , Mount Augustus National Park at 24°18.0ʹS 116°47.6ʹE (5 ♀, 4 ♂, USU) , at 24°19.2ʹS 116°48.9ʹE (1 ♀, ANIC; 1 ♀, CAS), and 24°22.8ʹS 116°54.2ʹE (1 ♀, ANIC) , 65 km E Nanutarra Roadhouse at 22°27.8ʹS 116°02.6ʹE (2 ♀, ANIC) , 158 km S Newman (= 9 km N Kumarina Roadhouse ) at 24°37.8ʹS 117°36.8ʹE [correctly: 119°36.8ʹE] (3 ♀, ANIC) , 24 km WNW Ophthalmia at 23°01.9ʹS 119°10.7ʹE (1 ♀, ANIC) , 47 km S Pardoo Roadhouse at 20°22.7ʹS 120°01.3ʹE (1 ♀, ANIC; 1 ♂, USU), 80 km S Pardoo Roadhouse at 20°28.3ʹS 120°10.0ʹE (1 ♀, CAS) , Perth: Darling Range (1 ♂, BMNH) , Perth: Darlington (3 ♀, WAM) , Perth: Kings Park (1 ♀, WAM) , 30 km ESE Three Rivers Station at 25°13.6ʹS 118°56.9ʹE (3 ♀, USU) , Turner Creek 24 km W Mulega junction 121 km W Highway 95 at 24°50.7ʹS 118°28.9ʹE (1 ♂, ANIC) , Walyunga National Park at 31°42ʹS 116°5ʹE (1 ♀, 1 ♂, CAS) .

NEW CALEDONIA: Noumea (Williams, 1945; Callan, 1960).

Figures

FIGURES 960-965. Pison rufipes Shuckard. (960) Female clypeus and mandibles; (961) Male clypeus and mandibles; (962) Female frons in lateral view showing erect setae; male: (963) Sternum VIII (ventral surface); (964) Genitalia in dorsal view; (965) Genitalia in lateral view.

FIGURES 80–85. Pison argentatum Shuckard. (80) Female clypeus and mandibles; (81) Male clypeus and mandibles; male: (82) Sternum VIII (ventral surface); (83) Genitalia in dorsal view; (84) Genitalia in lateral view; (85) Nest of Pison argentatum Shuckard.