Pison radians Pulawski, 2018

Pulawski, Wojciech J., 2018, A Revision of the Wasp Genus Pison Jurine, 1808 of Australia and New Zealand, New Guinea, and the Pacific Islands (Hymenoptera: Crabronidae), Proceedings of the California Academy of Sciences 65, pp. 1-584 : 386-388

Pison radians Pulawski , species nova

Figures 936 -941.

NAME DERIVATION.– Radians, present active participle of the Latin verb radiare, to radiate; with reference to the orientation of setae of the upper frons.

RECOGNITION.– The female of Pison radians (the male is unknown) is characterized by the presence of a psammophore on the mandible, lower gena ( Fig. 938 ), and forefemoral venter, and the lower gena unsculptured, shiny, and asetose between the hypostomal carina and the psammophore. It can be differentiated from the other species with these characters in having the setae characteristically radiating from the midpoint of the upper frons ( Fig. 937 ). The following combination is also diagnostic: gaster, femora and tibiae black; clypeal lamella nearly straight, with obtusely angular corner, minimally wider than distance that separates it from eye margin ( Fig. 936 ); mandible simple apically; ocellocular distance equal to 0.5-0.6 × hindocellar diameter, small- er than interocellar distance; scutal punctures nearly all less than one diameter apart, but not contiguous; most punctures of propleuron several diameters apart; propodeum with well-defined longitudinal carina separating side from dorsum and posterior surface and extending from gastral socket area toward spiracle; sterna punctate throughout; setae of frons, thorax, propodeal dorsum, and femora silvery, not completely concealing integument on mesopleuron and propodeal dorsum, but completely concealing on clypeus (except lamella); wings nearly hyaline; gastral terga with setal fasciae on apical depressions.

DESCRIPTION.– Frons dull, minutely punctate, punctures nearly contiguous. Occipital carina

Setae silvery, subappressed on upper frons, appressed on scutum, propodeal dorsum, and tergum I; radiating from midpoint on upper frons ( Fig. 937 ); completely concealing integument on clypeus; setal gena: see below. Apical depressions of terga with setal fasciae, fasciae silvery or with golden tinge.

Body black, mandible yellowish brown in basal half, apical depressions of terga brown, tarsi ferruginous.

♀.– Upper interocular distance equal to 0.68-0.70 × lower interocular distance; ocellocular distance equal to 0.5-0.6 × hindocellar diameter, distance between hindocelli equal to 0.8-1.2 × hindocellar diameter; eye height equal to 0.94-0.98 × distance between eye notches. Free margin of clypeal lamella nearly truncate, with well-defined lateral corner ( Fig. 936 ); distance between corners 1.1-1.2 × distance between corner and adjacent orbit. Dorsal length of flagellomere I 1.5-1.7 × apical width, of flagellomere IX 0.9-1.0 × apical width. Lower gena ( Fig. 938 ), mandibular posterior margin, propleural outer margin, and forefemoral venter ( Fig. 940 ) with psammophores (longest setae of genal, mandibular, and forefemoral psammophores about 0.7 ×, 0.7-1.0 ×, and 0.8 ×, respectively, of greatest forefemoral width); lower gena impunctate and asetose between hypostomal carina and psammophore. Mandible: trimmal carina with small incision at about midlength. Length 5.6-7.7 mm; head width 1.8-2.2 mm.

♂.– Unknown.

GEOGRAPHIC DISTRIBUTION (Fig. 941).–

Northern part of Northern Territory.

RECORDS.– HOLOTYPE: ♀, AUSTRALIA: Northern Territory: Keep River National Park at

15°57ʹ55ʺS 129°01ʹ52ʺE, 13-20 June 2001, M.E.

Irwin, F.D. Parker, and C. Lambkin (ANIC).

PARATYPES: AUSTRALIA: Northern Territory :

Keep River National Park at 15°57ʹ33ʺS

129°01ʹ44ʺE, 1 May – 3 June 2001, T. Weir,

K. Pullen, and P. Bouchard (1 ♀, CAS); Victoria

Highway 38.5 km SW Timber Creek at 15°42ʹ40ʺS

130°07ʹ48ʺE, M.E. Irwin, F.D. Parker, and C. Lambkin, 6-13 June 2001 (1 ♀, CAS), 13-19 June 2001

(2 ♀, CAS), 15-19 June 2001 (1 ♀, ANIC; 1 ♀, FIGURE 941. Collecting localities of Pison radians CAS ). Pulawski, sp. nov .

Figures

FIGURES 936-940. Pison radians Pulawski, sp. nov., female. (936) Clypeus and mandibles; (937) Upper frons; (938) Lower gena in lateral view showing psammophore; (939) Head in dorsal view; (940) Forefemur showing psam- mophore.