Pison stenometopon Pulawski, 2018

Pulawski, Wojciech J., 2018, A Revision of the Wasp Genus Pison Jurine, 1808 of Australia and New Zealand, New Guinea, and the Pacific Islands (Hymenoptera: Crabronidae), Proceedings of the California Academy of Sciences 65, pp. 1-584 : 435-438

Pison stenometopon Pulawski , species nova

Figures 1056-1062.

NAME DERIVATION.– Stenometopon derives from two Greek words: στενός, narrow, and μέτωπΟν, forehead, front; with reference to the unusually narrow upper interocular distance of this species female. RECOGNITION.– Pison stenometopon is an all black species, with three submarginal cells, the second recurrent vein ending at the very apex of the second submarginal cell, the setae appressed on tergum I, the tegula finely punctate throughout, and a carina separaing the propodeal side from the dorsum and the posterior surface. The female is characterized by the presence of a psammophore on the lower gena and the gena unsculptured and asetose between the hypostomal carina and the psammophore. It differs from similar species by the following combination: body all black, tegula finely punctate throughout, genal psammophore short (setal length equal to midocellar diameter), and forefemur without a psammophore (its longest setae shorter than midocellar diameter). In addition, the clypeal lamella is widely arcuate and forms an obtuse angle on each side, the distance between the angles being equal to the distance between an angle and the adjacent orbit.

The male can be recognized by its all black body, the tegula finely punctate throughout, and sternum VIII rounded apically. The cylindrical flagellum and the presence of short erect setae on sterna V-VII (setal length up to 0.7 × midocellar diameter) are subsidiary recognition features.

DESCRIPTION.– Frons dull, minutely punctate, punctures less than one diameter apart. Distance between antennal socket and orbit about equal to socket width. Gena narrow in dorsal view (Fig. 1057). Labrum not emarginate. Anteromedian pronotal pit transversely elongate, about as long as midocellar diameter. Propleuron largely impunctate, minutely microsculptured. Scutum finely foveate along flange, with or without short longitudinal ridges adjacent to posterior margin; scutal punctures fine, less than one diameter apart; interspaces finely microsculptured (Fig. 1058). Scutellum foveate along anterior margin. Tegula slightly enlarged, finely punctate throughout. Mesopleural punctures fine, less than one diameter apart. Postspiracular carina present, slightly shorter than midocellar diameter. Metapleural sulcus somewhat costulate between dorsal and ventral metapleural pits. Propodeum with irregular longitudinal carina separating side from dorsum and posterior surface and extending from gastral socket area toward spiracle; dorsum obliquely ridged, ridges evanescent toward lateral and posterior margins; side ridged (ridges well defined dorsally, evanescent ventrally), punctate between ridges; posterior surface irregularly ridged, with several conspicuous ridges radiating up from transverse carina just above gastropropodeal articulation. Forewing with three submarginal cells; second recurrent vein ending at very apex of submarginal cell II. Hindcoxal dorsum with outer margin obtusely carinate. Outer surface of hindtibia with evanescent spines. Punctures of horizontal part of tergum I, anterior to apical depression, with minute punctures about one diameter apart. Sterna minutely punctate throughout.

Setae silvery, appressed on frons, postocellar area, scutum, and tergum I, fully appressed on mesopleuron, not concealing integument on clypeus (see below for genal setae). Apical depressions of terga with inconspicuous silvery setal fasciae.

Body all black.

♀.– Upper interocular distance equal to 0.46-0.58 × lower interocular distance; ocellocular distance equal to 0.2-0.6 × hindocellar diameter, distance between hindocelli equal to 0.8-1.0 × hindocellar diameter; eye height equal to 0.96 × distance between eye notches. Free margin of clypeal lamella broadly arcuate, forming obtuse angle at each side (Fig. 1056); distance between angles equal to distance between angle and adjacent orbit. Dorsal length of flagellomere I 2.0-2.3 × apical width, of flagellomere IX 1.1-1.2 × apical width. Lower gena with straight, erect setae forming short psammophore (setae of psammophore as long as midocellar diameter), shiny, unsculptured, asetose between hypostomal carina and psammophore. Mandible: trimmal carina with small incision shortly beyond midlength. Forefemoral venter without psammophore, its longest setae less than one midocellar diameter long. Length 5.1-7.0 mm; head width 1.7-2.4 mm.

♂.– Upper interocular distance equal to 0.76 × lower interocular distance; ocellocular distance equal to 1.0 × hindocellar diameter, distance between hindocelli equal to 1.4-1.5 × hindocellar diameter; eye height equal to 0.96 × distance between eye notches. Free margin of clypeal lamella acutely angulate. Dorsal length of flagellomere I 1.7-2.1 × apical width, of flagellomere X 1.0 × apical width. Sterna V-VII with short erect setae (setal length up to 0.7 × midocellar diameter); sternum VIII rounded apically (Fig. 1059). Genitalia: Figs. 1060, 1061. Length 6.0- 6.5 mm; head width 1.9-2.1 mm.

GEOGRAPHIC DISTRIBUTION (Fig. 1062).–

Northern Territory, South Australia, Western

Australia.

RECORDS.– HOLOTYPE: ♀ AUSTRALIA: Northern Territory: Ngarradj Warde, Djobkeng in

Kakadu National Park, 27 June 1980, I.D. Naumann

(ANIC).

PARATYPES: AUSTRALIA: South Australia :

Calperum Station 14 km WNW Renmark at 34°07ʹS

140°37ʹE, 7 Nov – 13 Dec 1995, K.R. Pullen (1 ♀,

CAS). Western Australia: Fitzgerald River National Park at 33.949416°S 119.926086°E, 17 Jan 2010 ,

S. Krause (2 ♀, CAS; 1 ♀, MNKB) and L. Breit-

kreutz and S. Krause, 16-18 Jan 2010 (1 ♂, MNKB)

and 18-22 Jan 2010 (1 ♂, CAS); Yalgorup National FIGURE 1062. Collecting localities of Pison stenometo- Park at 32.880160°S 115.682545°E, 27 Jan 2010, S. pon Pulawski, sp. nov .

Krause (1 ♀, MNKB) .