Pison glabrum Kohl, 1908

Pulawski, Wojciech J., 2018, A Revision of the Wasp Genus Pison Jurine, 1808 of Australia and New Zealand, New Guinea, and the Pacific Islands (Hymenoptera: Crabronidae), Proceedings of the California Academy of Sciences 65, pp. 1-584 : 534-537

Pison glabrum Kohl

Figures 1293-1303.

Pison glabrum Kohl, 1908:309 , ♀. Holotype by monotypy: ♀, Samoa: Upolu ( NHMW), examined. – Turner, 1916b:626 (diagnostic characters); Perkins and Cheesman, 1928:6 (listed from Samoa), 26 ( Samoa, description of ♂); Yasumatsu, 1953:134 (in list of Pison of Pacific Islands); R. Bohart and Menke, 1976:335 (in checklist of world Sphecidae ); Dollfuss, 1989:11 (type material in NHMW); nec Naumann, 1990a:24 and Smithers, 1998:46 (= Pison laeve ); Kami and Miller, 1998:57 (in checklist of Samoan insects).

STATUS OF TYPE MATERIAL. – In the original description, Kohl did not specify the number of specimens examined, but his expression “Bei dem einzigem Stück …” [= “in the only exemplar”] demonstrates that he had only one. This specimen, therefore, is the holotype by monotypy.

RECOGNITION. – Pison glabrum closely resembles P. laeve in having an unsculptured, sparsely punctate mesopleuron and propodeal dorsum and side (propodeum without the longitudinal carina separating the dorsum and side), and the ocellocular distance equal to 0.3 × hindocellar diameter. It differs from P. laeve in having the scutal flange the usual shape (not expanded), the posteroventral forefemoral punctures relatively well defined (rather than microscopically small), the microscopic punctures on the disk of terga I and II many diameters apart (rather than a few diameters), sternum II mostly or all impunctate (rather than sparsely punctate throughout), and female terga without silvery, apical fasciae (rather than fasciate).

Also similar is Pison insulare , from which P. glabrum differs in having the setae appressed on scutum and tergum I, impunctate apical depressions of terga I-IV, and in lacking silvery setae on terga. In P. insulare , the setae are erect on the scutum and in most specimens on the basal part of tergum I, the apical depressions of at least terga I and II are microscopically punctate and covered with silvery, setal fasciae.

DESCRIPTION.– Frons microareolate, slightly shiny, with well defined punctures, most punctures many diameters apart (Fig. 1294). Gena narrow in dorsal view. Labrum not emarginate. Anteromedian pronotal pit transversely elongate, about as long as midocellar diameter. Scutum not foveate along flange, without short longitudinal ridges adjacent to posterior margin; scutal punctures averaging several diameters apart, interspaces minutely microsculptured. Tegula enlarged. Mesopleural punctures in most specimens several diameters apart at center (Fig. 1295), about two diameters in specimen from Moorea Island. Postspiracular carina present, about twice as long as midocellar diameter. Metapleural sulcus conspicuously costulate between dorsal and ventral metapleural pits. Propodeum without longitudinal carina separating side from dorsum and posterior surface and extending from gastral socket area toward spiracle; dorsum with minute punctures that are many diameters apart, unsculptured between punctures, lateral punctures larger in holotype, averaging 2-3 diameters apart (Fig. 1296); side finely microsculptured, microscopically punctate, punctures many diameters apart, larger posteriorly, averaging 2-3 diameters apart (Fig. 1297); posterior surface punctate except ridged in ventral third. Posteroventral forefemoral surface microsculptured, with a few, sparse punctures. Hindcoxal dorsum with outer margin carinate in distal half. Punctures of tergum I microscopic, several diameters apart, apical depressions of terga I-V impunctate (except punctate laterally). Sterna II and III impunctate except for several punctures posterolaterally.

Setae suberect, oriented ventrad on upper frons (mostly about as long as midocellar diameter, uppermost setae longer than midocellar diameter); appressed on scutum and tergum I; erect, slightly sinuous on lower gena (length about 1.5 × midocellar diameter); not concealing integument on clypeus. Terga sparsely setose (practically asetose in female), apical depressions in female without setal fasciae, in male with evanescent fasciae.

Body all black, wing membrane infumate.

♀.– Upper interocular distance equal to 0.52 × lower interocular distance; ocellocular distance equal to 0.3 × hindocellar diameter, distance between hindocelli equal to 0.6 × hindocellar diameter; eye height equal to 1.04 × distance between eye notches. Free margin of clypeal lamella obtusely angulate (exactly as in P. laeve , see Fig. 560 ). Dorsal length of flagellomere I 2.4 × apical width, of flagellomere IX 1.5 × apical width. Mandible: trimmal carina with small incision shortly beyond midlength. Length 8.2-9.0 mm; head width 2.4-2.6 mm.

♂.– Upper interocular distance equal to 0.62 × lower interocular distance; ocellocular distance equal to 0.8 × hindocellar diameter, distance between hindocelli equal to 0.7 × hindocellar diameter; eye height equal to 1.10 × distance between eye notches. Free margin of clypeal lamella straight with rounded median point, slightly convex on each side of point, slightly concave laterally (Fig. 1293). Flagellomeres III-VIII slightly convex ventrally. Dorsal length of flagellomere I 2.4 × apical width, of flagellomere X 1.7 × apical width. Sternum VIII with prominence, shallowly, broadly emarginate apically (Fig. 1299), in oblique lateral view (Fig. 1300). Genitalia: Figs. 1301, 1302.

GEOGRAPHIC DISTRIBUTION (Fig. 1303).– Federated States of Micronesia, Samoa and also

Moorea and Austral Islands in French Polynesia.

RECORDS.– FEDERATED STATES OF MICRONE-

SIA: Pohnapei: Jokai Island (1 ♀, BISH) .

FRENCH POLYNESIA: Austral Islands: Rurutu

Island: Moerai (1 ♂, BISH). Moorea: base of Mount

Tohivea: Belvedere Trail (1 ♀, BISH, as Tohihea) .

SAMOA: no specific locality (1 ♀, NHMW,

holotype of Pison glabrum ). Savaii: no specific locality (Perkins and Cheesman, 1928). Tutuila:

Amouli (1 ♀, CAS), Aua-Afono Trail (2 ♀, BISH) ,

Fagasa (1 ♀, BISH), Leone Auila (1 ♀, BISH) ,

Leone Aulau Trail (1 ♀, BISH), no specific locality FIGURE 1303. Collecting localities of Pison glabrum (1 ♀, CAS). Upolu: Mount Vaea (1 ♂, BISH), Tuae- Kohl .

fu (1 ♀, 1 ♂, BMNH) .

Figures

Figures 560-565. Pison laeve F. Smith. (560) Female clypeus and mandibles; (561) Male clypeus and mandibles; (562) Female vertex; (563) Left part of female scutum (arrow shows expanded flange); (564) Female mesopleuron; (565) Propodeal dorsum of female.