Empis doronicola, 2012

Çiftçi, Mustafa Cemal, Hasbenli, Abdullah & Canpolat, Derya, 2012, Two new species of Empis (Diptera: Empididae) from Turkey, Zootaxa 3408, pp. 47-53 : 50

Empis (s.str) doronicola Çiftçi sp. nov.

( Figs. 4–6 )

Diagnosis. Large (body 6–7 mm long), thoracic and abdominal hairs and bristles whitish yellow, legs and upper occipital hairs black. Halteres yellow with darker stem. Female legs without pennation or flattened hairs.

Description. Male. Head black with grey dusting. Eyes with upper ommatidia enlarged. Face and frons grey dusted, face wide and broadened through mouth edge. Occiput grey, upper half of occiput with 2 rows of black occipital bristly-hairs, lower half around mouth opening with yellowish hairs. Antenna black, 1st (scape) and 2nd (pedicel) antennal segment with short black hairs, stylus as long as 3rd antennal segment (postpedicel). Palpus black, with thin black hairs. Labrum twice as long as height of head.

Thorax densely grey dusted, scutum also with slightly yellowish dusting. Scutum in frontal and dorsal view subshinning, in posterior and lateral view with 3 dull black stripes. Mesonotum densely clothed with yellowish hairs leaving only area between acrostichal and dorsocentral rows bare. Acrostichals and dorsocentrals yellowish, nearly as long as antennal stylus, acrostichals irregularly 3–4 serial, multiserial dorsocentral ending with 3–4 long yellowish prescutellar pairs. Large thoracic bristles yellowish, humeral bristle not differentiated because of long hairs on humeri (postpronotum), 4 notopleural, 2 supra-alars, 1 postalar and 2 pairs of scutellar yellowish and distinct. Prothorax with yellowish white hairs, pronotum with row of short hairs. Metapleural fan (laterotergal bristles) whitish.

Wing very faintly brownish, all veins dark and distinct, only wing base yellowish. Costal bristle not differentiated, stigma black. Anal vein complete. Squama yellowish with pale fringes. Haltere yellow with darker stem.

Legs long and slender, shinning black except for densely grey coxae. Hind tibia dilated towards tip, fore basitarsus slightly, hind basitarsus clearly swollen but not wider than tip of hind tibia. Coxae with whitish hairs. Fore femur dorsally with black, ventrally with sparse long yellowish hairs. Mid and hind femora with black dorsal hairs on apical half, other hairs pale, basally long shortened towards tip. Ventral hairs on mid and hind femora bristlelike. Hind femur anteroventrally with row of bristly hairs slightly longer than width of hind femora. Fore tibia slen- der, covered with short black hairs, dorsal hairs slightly longer. Mid tibia ( Fig. 4 ) with short black hairs, anterodorsally and ventrally with row of bristles, apically and basally shortened ventral row long at middle. Hind tibia dorsally with 2 rows of bristles, ventrally with short adpressed bristly hairs. Tarsal segments with short hairs and apical circlets of bristly-hairs. Fore and mid basitarsi and hind tarsal segments ventrally with short spine-like hairs. Hind basitarsus dorsally with long hairs.

Abdomen black, dorsum black, subshinning and slightly grey dusted, sterna densely grey dusted. Abdominal hairs whitish, long on anterior segments and shortened towards tip. Sternum 8 with long yellowish bristles on hind margin. Genitalia (terminalia) ( Fig. 5 ) large and distinct. Aedeagus (phallus) ( Fig. 6 ) brownish yellow, long and bow-shaped, sligthly thickened at base. Lateral lamella (epandrium) triangular, hind margin with long pale hairs. Dorsal lamella (cercus) with short dark hairs, anteriorly oval, suddenly narrowed and finger-like at tip.

Holotype male body length: 6.2 mm, wing length: 6.8 mm. Paratypes body length: male 5.6–6.4 mm; female 6.5–7.1 mm, wing length: male 6.2–7 mm; female 6.3–6.9 mm.

Female. Main differential characters and general appearance as in male with shorter hairs. Wings distinctly brownish. Abdomen densely grey dusting on terga and sterna. Legs shinning black but especially on femora with slightly grey dusting. Fore basitarsus (tarsomere 1) simple, hind basitarsus and tip of hind tibia little swollen as in male. Legs without pennation and flattened hairs. Other characters on legs as in male with shorter bristles.

Remarks. Considering the dense, long and soft yellowish white hairs on the mesonotum, E. doronicola is very much like E. decora Meigen , but in the former species the mesonotum has only yellowish white hairs including strong notopleurals, supra-alars, postalar and also scutellar setae. Empis decora have black bristling and hairing on posterior part of mesonotum. Female legs not pennate in E. doronicola but in E. decora female legs are pennate.

Type material. HOLOTYPE 3 (spn13760): TURKEY. Yozgat, Akdaġmadeni, north of Davulbaz village, 1737 m, 06.vi.2011. PARATYPES (spn13761-13802): TURKEY. 83, 10Ƥ, Yozgat, Akdaġmadeni, SE of Özer village, 1839 m, 06.vi.2011; 133, 11Ƥ, Yozgat, Akdaġmadeni, N of Davulbaz village, 1737 m, 06.vi.2011.

Derivatio nominis. The species is named after the plant Doronicum orientale Hoffmann, 1808 from the family Asteraceae , upon which the type material were collected.

Figures

FIGURES 4 – 6. Empis (s. str.) doronicola 4. Mid leg in posterior view; 5. Postabdomen; 6. Aedeagus (phallus). Scale = 0.3 mm.