Montiphylax antennatus (Banks, 1900)

Ruiter, David E. & Mutch, Robert A., 2019, Montiphylax, (Trichoptera, Limnephilidae), a new genus to accommodate the western North American species: Stenophylaxantennatus Banks, 1900, Philocascathor Nimmo, 1971, and Philocascaalba Nimmo, 1977, ZooKeys 845, pp. 153-180 : 157-159

Montiphylax antennatus (Banks, 1900) Figures 1, 4C, D, 6, 7

Stenophylax antennatus Banks, 1900: 254-255 (male description) Washington.

Anisogamus antennatus Milne, 1935: 29; Banks, 1943: 350, fig. 41 and 50 (describes 2nd specimen from Wallace, Idaho. This Idaho specimen is M. thor , see discussion below.).

Stenophylax antennatus Ross, 1944: 299.

Philocasca antennata Schmid, 1955: 201; Flint, 1966: 379, fig. 2m, n; Wiggins & Anderson, 1968: 74; Wiggins, 1977: 274; Mutch, 1981: 223; Wiggins, 1996: 338; Schmid, 1998: 121; Ruiter et al., 2005: 162; Blinn & Ruiter, 2013: 292.

Description.

Head to wingtip length: male 19-21 mm (n = 5); female 19-21 mm (n = 5).

Male genitalia (Fig. 6): 8th tergite without dorsal modified spines or projections. Ninth segment annular with tergite narrow and strap-like, directed distad dorsally resulting in slightly sigmoid anterior margin of 9th in lateral view; remainder of 9th broad in lateral view, directed downward at approximately a 45 degree angle from the narrow tergite; inferior appendages appearing separated from 9th. In caudal view, inferior appendages slightly cupped around phallic apparatus and slightly separated mesally. Tenth segment cupped anteriorly, appearing as two, slightly fused hemispheres extending broadly anteriad within the 9th. Superior appendage nearly twice as long as tall in lateral view. Intermediate appendages arise ventrally, strongly sclerotized and extending as two long, tapered, parallel projections, curving downward at apex. In caudal view, below the intermediate appendages are paired, curved, narrow projections nearly surrounding the anus. Phallus large, with strongly sclerotized phallicata; membranous endophallus with dorsal, strongly sclerotized band, projected upward and distad apically; strong, thick parameres originate at the base of the dorsal band, extend distally; tapering to blunt, rounded apex in lateral view and slight curved upward throughout. Parameres originate dorsolaterally and extend posteriorly over 3 times as long as wide; thick throughout, ending in blunt, rounded apex in dorsal view. The aedeagus apex a sclerotized tube within an extensile sheath; ending in the bottom of a posteriorly directed sclerotized cup.

Allotype female genitalia (Fig. 7): 9th segment fused laterally and incomplete ventrally, separated ventrally by broad supragenital plate. Ninth fused with 10th; 9th tergite ca. half as long as 10th in lateral view; in lateral view 10th with dorsal and ventral margins slightly tapered throughout to rounded apex. In dorsal view, 10th cleft less than half the distance to base. In lateral view, ventrolateral corners of 9th extend distally beyond remainder of 9th; acute at apex and directed inward in ventral view. The medial lobe of the vulval scale short, shorter than lateral lobes, longer than wide. The vaginal apparatus, in lateral view, rectangular, ca. twice as long as tall; in ventral view, egg-shaped with smoothly rounded anterior margin.

Material examined.

WA: Whatcom County, stream near Heather Meadow Information Center on Artist Point Road, D.W. Blinn, 28 July 2009, blacklight trap, 2M; stream on Fire and Ice Trail, near top of Mt. Baker Highway, 48.85586 -121.68795, D.W. Blinn, 13 August 2012, 2L, 1M, 1F; spring stream, ca. 150 feet southeast of outhouse at Mt. Baker Information Center parking lot, near top of Highway 542, 48.85378 -121.68543, D.E. Ruiter, 5 August 2012, 1L; meltwater stream on Fire and Ice trail, in Heather Meadows, Mt. Baker, D.W. Blinn, 31 July 2009, 13M, 3F (2M to National Museum of Natural History; 2M, 1F to S. Chuluunbat; 1M, 1F to Canadian National Collection, remainder in D.E. Ruiter personal collection); Terminal Lake on Mt. Baker Highway, D. W. Blinn, black light trap, 28 July 2009, 1M.