Coniceromyia maculata, Ament & Kung & Brown, 2020

Ament, Danilo César, Kung, Giar-Ann & Brown, Brian V., 2020, Forty-one new species of Coniceromyia Borgmeier (Diptera: Phoridae), an identification key, and new distributional records for the species of the genus, Zootaxa 4830 (1), pp. 1-61 : 27-28

Coniceromyia maculata sp. nov.

( Figs. 4E , 8W , 14K , 20 G–H, 24X, 27M)

Diagnosis (male). Foremetatarsus widening towards apex, highly setulose, with anterior excavation, broad and short anteroapical expansion, without differentiated long setae, and basal half dark brown ( Fig. 8W ).

Material examined. Holotype ♂, COLOMBIA: Boyacá: SFF Iguaque : Cab. Mamarramos, 5.69ºN, 76.45ºW, 8–25.vi.2000, Malaise trap, 2855m, P. Reina col. ( IAVH) [LACM ENT 211148] . Paratypes: 1♂, Boyacá: SFF Iguaque : La Planada, 5.42ºN, 73.45ºW, 23.ix–11.x.2000 , Malaise trap, 2850m, P. Reina col.; 1♂, same but 30.vii– 17.viii.2000; 2♂, same but 5.70ºN, 73.46ºW, 28.ii–16.iii.2000; 2♂, same but 1–19.iv.2000; 1♂, same but 5.42ºN, 73.45ºW, 14.iv–6.v.2000; 1♂, Santander: Gambitá : Vda. El Palmar: Sector el Taladro, 05.98°N, 73.22°W, 14– 31.v.2018 , Malaise trap, 2357m, T. Zambrano-Estupiñan ( CEUA) .

Description. Male. Body length, 2.5 mm. Head. Frons brown, shining, sparsely setulose, with well-defined median furrow. Flagellomere 1 light brown, conical very elongate. Arista apical, about one-third flagellomere 1 length. Palpus yellowish-brown. Gena with two setae. Thorax. Scutum, scutellum, and pleural regions brown, shining; anepisternum bare. Legs yellowish-brown, except foremetatarsus basal half dark brown, hind tibia and hind tarsus light brown, and hind femur apex dark brown. Forefemur with ventral row of slightly elongate setae and dorsal rounded macula near apex. Foretibia with two dorsal setae and anterodorsal row of strong setulae. Foremetatarsus widening towards apex, highly setulose, with anterior excavation, broad and short anteroapical expansion, without differentiated long setae ( Fig. 8W ). Foremetatarsus ratio 0.35. Hind femur expanded ventrally, with broad basoventral band of relatively long setulae ( Figs. 20 G–H). Hind femur height/length ratio 0.42. Hind tibia with one dorsal seta in basal half. Wing ( Fig. 14K ). Costa considerably swollen, 0.48 of the wing length; R 2+3 absent; M 1 concave, curved posteriorly at apex; M 2 basal half straight, apical half sinuous, CuA 1 -M 2 /M 2 -M 1 ratio 1.5; CuA 1 strongly sinuous. Halter yellowish-white. Abdomen. Tergites dark brown, with shining pubescence. Hypopygium light brown ( Fig. 24X ), left epandrial process with a subepandrial wall extending from the hypoproct to its apex and a subepandrial group of strong setulae. Margin of right lobe of hypandrium without lateral projection. Hypoproct with three setae.

Distribution. High-elevation sites in Colombian Eastern Andean Cordillera ( Fig. 27M ).

Etymology. Name derived from the Latin root macula (spot, mark), referring to the macula on the base of the foremetatarsus.

Variations. Specimens with a conspicuous basoventral macula on the hind femur posterior face.

Figures

FIGURE 4. Habitus of the holotypes of new species of Coniceromyia, continuation (scale bars=1mm). A. C. inflata, B. C. leia, C. C. litopoda, D. C. luna, E. C. maculata, F. C. megalosoma.

FIGURE 8. Male foremetatarsus of new species of Coniceromyia (scale bars=0.1mm). A. C. amazonensis, B. C. anisocrossa, C. C. balbiae, D. C. bellatula, E. C. bisetosa, F. C. brachypoda, G. C. camptoneura, H. C. chrysopa, I. C. circulata, J. C. cladopyga, K. C. crassivena, L. C. dasypoda, M. C. deltopoda, N. C. diadela, O. C. diffusa, P. C. dolichopoda, Q. C. hadrochaeta, R. C. hirtipenna, S. C. inflata, T. C. leia, U. C. litopoda, V. C. luna, W. C. maculata, X. C. megalosoma, Y. C. memorialis, Z. C. niemeyeri, A’. C. numerosa, B’. C. ochyrosticha, C’. C. parallela, D’. C. pectinella.

FIGURE 14. Wings of the males of Coniceromyia new species, continuation. A. C. deltopoda, B. C. diadela, C. C. diffusa, D. C. dolichopoda, E. C. hadrochaeta, F. C. hirtipenna, G. C. inflata, H. C. leia, I. C. litopoda, J. C. luna, K. C. maculata, L. C. megalosoma.

FIGURE 20. Hind femora of the males of Coniceromyia new species, continuation (scale bars on figures of the entire femur=0.3 mm, on figures of ventrobasal part of the femur=0.1 mm). A–B. C. leia, C–D. C. litopoda, E–F. C. luna, G–H. C. maculata, I. C. megalosoma, J. C. memorialis

FIGURE 24. Hypopygia of Coniceromyia new species in right lateral view (except figure J which is left lateral view). A. C. amazonensis, B. C. anisocrossa, C. C. balbiae, D. C. bellatula, E. C. bisetosa, F. C. brachypoda, G. C. camptoneura, H. C. chrysopa, I. C. circulata, J. C. cladopyga in left lateral view, K. C. cladopyga in right lateral view, L. C. crassivena, M. C. dasypoda, N. C. deltopoda, O. C. diadela, P. C. diffusa, Q. C. dolichopoda, R. C. hadrochaeta, S. C. hirtipenna, T. C. inflata, U. C. leia, V. C. litopoda, W. C. luna, X. C. maculata.

FIGURE 27. Distributional records of Coniceromyia new species (species known from a single or close localities were not illustrated. Squares=holotype, circles=other specimens). A. C. amazonensis, B. C. bellatula, C. C. bisetosa, D. C. brachypoda, E. C. cladopyga, F. C. dasypoda, G. C. deltopoda, H. C. diffusa, I. C. hadrochaeta, J. C. hirtipenna, K. C. litopoda, L. C. luna, M. C. maculata.