Ischnothyreus culleni, Edward & Harvey, 2014

Edward, Karen L. & Harvey, Mark S., 2014, Australian Goblin Spiders Of The Genus Ischnothyreus (Araneae, Oonopidae), Bulletin of the American Museum of Natural History 2014 (389), pp. 1-144 : 26-29

Ischnothyreus culleni , new species Figures 9–10 , map 4

TYPES: AUSTRALIA: Queensland: Male holotype and female allotype from sifted leaf leater in Mount Cook National Park off walking track, near lookout, 15.47444 ° S, 145.26361 ° E (12 May 2008, K. Edward and P. Cullen), deposited in QM (³ holotype: QM S95902 , PBI _ OON 00005612 ; ♀ allotype: QM S95903 , PBI _ OON 00005539 ) .

ETYMOLOGY: The specific name is a patronym in honor of Patrick Cullen, one of the collectors of the holotype and many other interesting creatures, and in recognition of his passion for the natural world.

DIAGNOSIS: Females of I. culleni , sp. nov., are quite similar to I. nourlangie , sp. nov. (fig. 6F), and I. monteithi , sp. nov. (fig. 59F), as they all have a very simple epigynal region, with no discerning epigynal atrium. Howev- er, females of this species can be distinguished by a yellow–pale orange colored carapace, a short dorsal scute, covering 1/2 of the abdomen length, and a small triangular process at posterior end of a convoluted duct (figs. 9D, I, 10F). The male pedipalps possess an enlarged clublike embolic region, and the dorsal surface of the fang has a sharp triangular incision that is not heavily sclerotized (fig. 10A–D).

MALE (PBI_OON 5612, figs. 9A–C, G–H, 10A–E). Total length 1.20. CEPHALOTHO- RAX: Carapace pale orange, ovoid in dorsal view, pars cephalica strongly elevated in lateral view, anteriorly narrowed to 0.49 times its maximum width or less, anterolateral corners without extension or projections, surface of elevated portion of pars cephalica smooth, sides finely reticulate; nonmarginal pars cephalica setae dark. Clypeus margin unmodified, curved downward in front view, high, ALE separated from edge of carapace by their radius or more; setae dark. Eyes: ALE largest, ALE circular, PME oval, PLE oval; posterior eye row procurved from above; ALE touching, ALE-PLE touching. Sternum longer than wide, yellow, uniform; setae dark, evenly scattered. Chelicerae slightly divergent, anterior face unmodified; promargin with one larger denticle; fang shape normal, without prominent basal process; fang modified with triangular indent subbasally; setae dark. Labium elongated hexagon, not fused to sternum, anterior margin slightly indented at middle, much more heavily sclerotized than sternum; with six or more setae on anterior margin, subdistal portion with unmodified setae. Endites anteromedian tip with one strong, toothlike projection, much more heavily sclerotized than sternum. ABDOMEN: Ovoid; dorsum soft portions white. Book lung covers elliptical. Dorsal scutum pale orange, covering more than 3/4 of abdomen, more than 1/2 to most of abdomen width, middle surface smooth, sides smooth. Postepigastric scutum pale orange, covering about 1/2 of abdominal length. Epigastric area and postepigastric area setae dark. LEGS: Yellow, without color pattern; patella plus tibia I shorter than carapace. Leg spination: femora: I p0-1-1; II p0-1-0; tibiae: I, II p2-2-0; v2-2-0; metatarsi: I, II p1-1-0; v1- 1-0. GENITALIA: Palp proximal segments dark red-brown; embolus dark; femur shorter than trochanter, with posteriorly rounded lateral dilation; patella about as long as femur; cymbium dark red-brown, without distal patch of setae; bulb dark red-brown, more than two times as long as cymbium, stout; embolic region, stout, enlarged, clublike (fig. 10A, B, E).

FEMALE (PBI_OON 5539, figs. 9D–F, I, 10F). Total length 1.65. CEPHALOTHO- RAX: Carapace yellow, pars cephalica slightly elevated in lateral view. Clypeus low, ALE separated from edge of carapace by less than their radius. Chelicerae, endites, and labium yellow. Dorsal scutum covering about 1/2 of abdomen, between 1/4 and 1/2 abdomen width. Epigastric scutum small lateral sclerites present. Postepigastric scutum widely hexagonal, covering about 1/3 of the abdominal length. LEGS: Pale orange. Leg spination: femora: I p0-1-1; II p0-1-0; tibiae: I, II p2-2-0; v2-2-0; metatarsi: I, II p1-1-0; v1-1-0. GENITALIA: Ventral view: epigynal region very simple, without distinct epigynal atrium; convoluted duct ends at small posteriorly pointed triangular process; convoluted duct thicker than apodemes (fig. 10F).

OTHER MATERIAL EXAMINED: AUS- TRALIA: Queensland: Black Mountain National Park , 15.68333 ° S, 145.21666 ° E, July 20–Nov. 28, 1992 (R. Raven, P. Lawless, M. Shaw, QM S24151 , PBI _OON 21994), 1 ³ ;

Black Mountain National Park , across road from Black Mountain lookout, vine thicket and eucalypt among boulders, leaf litter, 15.66083 ° S, 145.22111 ° E, May 12, 2009 (K. Edward and P. Cullen, QM S95904 ), 1 ³, 4 ♀ ; Black Mountain National Park , across Road from Black Mountain lookout, vine thicket and eucalypt among boulders, leaf litter, 15.66083 ° S, 145.22111 ° E, May 12, 2009 (K. Edward and P. Cullen, WAM T130759 , PBI _ OON 5541 ), 1 ³, 1 ♀ ; Black Mountain National Park , across Road from Black Mountain lookout, vine thicket and eucalypt among boulders, leaf litter, 15.66083 ° S, 145.22111 ° E, May 12, 2009 (K. Edward and P. Cullen, QM S95905 , PBI _ OON 5574 ), 1 ♀ ; Mount Cook National Park ; walking track, near lookout, rainforest, leaf litter amongst boulders, 15.47444 ° S, 145.26722 ° E, May 12, 2009 (K. Edward and P. Cullen, QM S95906 , PBI _ OON 5573 ), 1 ♀ ; Mount Cook National Park ; walking track, near lookout, rainforest, leaf litter amongst boulders, 15.47444 ° S, 145.26722 ° E, May 12, 2009 (K. Edward and P. Cullen, QM S95907 , PBI _ OON 5588 ), 1 ³, 2 ♀ ; Mount Cook National Park , 80 m, 15.50000 ° S, 145.26670 ° E, Nov. 10–16, 1975 (R. Raven, V. Davies, QM S16138 , PBI _OON 26244), 10 ³ ; Mount Cook , Quarantine Road (NQ 21), 15.50000 ° S, 145.23330 ° E, July 20– Nov. 28, 1992 (R. Raven, P. Lawless, M. Shaw, QM S24419 , PBI _OON 21802), 8 ³, 2 ♀ ; Shiptons Flat , 15.78333 ° S, 145.23333 ° E, Oct. 17–19, 1980 (T. Weir, ANIC, PBI _OON 25820), 1 ³, 1 ♀ ; Shiptons Flat Road , Rossville, 9 km S Helenvale, rainforest, leaf litter, 15.77944 ° S, 145.22750 ° E, May 12, 2009 (K. Edward and P. Cullen, QM S95908 ), 2 ³, 1 ♀ ; Shiptons Flat Road , Rossville, 9 km S Helenvale, rainforest, leaf litter, 15.77944 ° S, 145.22750 ° E, May 12, 2009 (K. Edward and P. Cullen, QM S95909 , PBI _ OON 5542 ), 1 ♀ ; Shiptons Flat Road , Rossville, 9 km S Helenvale, rainforest, leaf litter, 15.77944 ° S, 145.22750 ° E, May 12, 2009 (K. Edward and P. Cullen, QM S95910 , PBI _ OON 5575 ), 1 ♀ .

DISTRIBUTION: known only from three northern subregions of the Wet Tropics Bioregion (CL, BL, and FU), in northeastern Queensland (map 4).

Figures

Fig. 9. Ischnothyreus culleni, sp. nov. Holotype male (PBI_OON 00005612): A. habitus, dorsal view; B. habitus, ventral view; C. habitus, lateral view; G. carapace, anterior view; H. sternum, ventral view. Allotype female (PBI_OON 00005539): D. habitus, dorsal view; E. habitus, ventral view; F. habitus, lateral view; I. epigynum, ventral view.

Fig. 10. Ischnothyreus culleni, sp. nov. Holotype male (PBI_OON 00005612): A. left palp, prolateral view; B. left palp, retrolateral view; C. left chelicerae, anterior view; D. left chelicerae, posterior view; E. left palp, dorsal view. Allotype female (PBI_OON 00005539): F. epigynum, ventral view. Scale lines 5 0.1 mm.