Ischnothyreus hoplophorus, Edward & Harvey, 2014

Edward, Karen L. & Harvey, Mark S., 2014, Australian Goblin Spiders Of The Genus Ischnothyreus (Araneae, Oonopidae), Bulletin of the American Museum of Natural History 2014 (389), pp. 1-144 : 48-50

Ischnothyreus hoplophorus , new species Figures 23–24 , map 4

TYPES: AUSTRALIA: Queensland: Male holotype and female allotype from Mount Lewis , 16.35 ° S, 145.17 ° E, 960 m (30 Oct 1976, R.W. Taylor and A. Weir), deposited in ANIC (³ holotype: PBI _ OON 00005845 ; ♀ allotype: PBI _ OON 00005598 ) .

ETYMOLOGY: The specific epithet is derived from the Greek hoplon (‘‘armor, shield’’), meaning ‘‘bearing a shield’’ ( Brown, 1956), and refers to the distinctive heavily sclerotized plate of the female epigyinum region of this species.

DIAGNOSIS: This species is closely related to I. bifidus , sp. nov., as males of both species possess a clypeus that is concave medially (figs. 17H, 23G), the anterior face of the chelicerae is modified with complex processes (figs. 22C, 24C), and the hyaline grooves, anterolateral corners of the carapace, endites, and the lateral edges of the labium are all heavily sclerotized (fig. 17G–I, 23G–H). Females share a unique epigynal region that consists of a heavily sclerotized platelike structure anterior to an ovoid shaped epigynal atrium (figs. 22E, 24E). Ischnothyreus hoplophorus , sp. nov., can be distinguished by the nonbifurcated anterior process on the male chelicerae (fig. 24C), and the elongate platelike sclerite of the female epigynal region, which has curved lateral edges (figs. 23I, 24E).

MALE (PBI_OON 00005845, figs. 23A–C, G–H, 24A–D). Total length 1.58. CEPHA- LOTHORAX: Carapace pale orange, without any pattern, broadly oval in dorsal view, pars cephalica strongly elevated in lateral view, anteriorly narrowed to between 0.5 and 0.75 times its maximum width, anterolateral corners with strongly sclerotized, triangular extension, strongly sclerotized lines extend either side of clypeus half way to elevated portion of pars cephalica; surface of elevated portion of pars cephalica smooth, sides finely reticulate. Clypeus margin unmodified, very strongly curved in front view, low, ALE separated from edge of carapace by less than their radius; setae dark. Eyes: ALE largest, ALE circular, PME oval, PLE oval; posterior eye row straight from above; ALE separated by less than their radius, ALE-PLE touching. Sternum longer than wide, pale orange, uniform; setae dark, evenly scattered. Chelicerae, endites, and labium dark red-brown. Chelicerae slightly divergent, anterior face with hyaline grooves and distal complex projections, anterior face with three major heavily sclerotized projections, largest digitiform; fang shape normal, with prominent basal process, broad, fluted outgrowth; setae dark (fig. 24C, D). Labium elongated hexagon, not fused to sternum, anterior margin indented at middle, more sclerotized than sternum; with six or more setae on anterior margin. Endites anteromedian tip with one strong, toothlike projection, much more heavily sclerotized than sternum. ABDO- MEN: Ovoid; dorsum soft portions white. Book lung covers elliptical. Dorsal scutum pale orange, covering more than 3/4 of abdomen, more than 1/2 to most of abdomen width, middle surface smooth, sides smooth. Epigastric scutum small lateral sclerites present. Postepigastric scutum pale orange, covering about 2/3 of abdominal length. Dorsum, epigastric area, and postepigastric area setae dark. LEGS: Yellow, without color pattern; patella plus tibia I shorter than carapace. Leg spination: femora: I p0-1-1; II p0-1-0; tibiae: I, II p2-2-0; v2-2-0; metatarsi: I, II p1-1-0; v1-1-0. GENITALIA: Palp proximal segments dark red-brown; embolus dark; femur one to two times as long as trochanter; patella shorter than femur; cymbium dark red-brown; bulb dark red-brown, more than 2 times as long as cymbium, stout; embolic region obtusely bent, elongate, basal section broad, narrow distally, slightly curved distal tip; fine denticles present on ventral subdistal edge (fig. 24A, B).

FEMALE (PBI_OON 5598, figs. 23 D–F, I, 24E). Total length 1.90. CEPHALOTHO- RAX: Carapace pars cephalica slightly elevated in lateral view, anteriorly narrowed to 0.49 times its maximum width or less. Clypeus straight in front view. Eyes: ALE touching. Sternum orange-brown. Mouthparts: Chelicerae, endites, and labium pale orange. ABDOMEN: Dorsal scutum orangebrown, covering 1/2 to 3/4 of abdomen, between 1/4 and 1/2 abdomen width. Postepigastric scutum widely hexagonal, only around epigastric furrow. LEGS: Orangebrown. Leg spination: femora: I p0-1-1; II p0-1-0; tibiae: I, II p2-2-0; v2-2-0; metatarsi: I, II p1-1-0; v1-1-0. GENITALIA: Ventral view: anterior edge of epigastric furrow strongly procurved; epigynal region with heavily sclerotized platelike structure, elongate, lateral edges curved and undulate, epigynal atrium ovoid; convoluted duct thicker than apodemes, elongate (fig. 23I).

OTHER MATERIAL EXAMINED: AUS- TRALIA: Queensland: Mount Lewis , rainforest, leaf litter, 970 m, 16.33000 ° S, 145.13000 ° E, Sept. 8, 1975 (R.W. Taylor, ANIC, PBI _OON 25870), 1 ♀ ; Mount Lewis , 3 km E, 680 m, 16.58367 ° S, 145.30220 ° E, Nov. 29, 1997, to Feb. 09, 1998 (G. Monteith, D. Cook, QM S69564, PBI _OON 22174), 1 ³ ; Mount Lewis , barracks, 950 m, 16.61167 ° S, 145.27500 ° E, Nov. 17, 1997 (G. Monteith, QM S43210 , PBI _OON 21788), 1 ♀ .

DISTRIBUTION: This species is only known from Mount Lewis, northeastern Queensland (map 4; CU subregion).

Figures

Fig. 23. Ischnothyreus hoplophorus, sp. nov. Holotype male (PBI_OON 00005845): A. habitus, dorsal view; B. habitus, ventral view; C. habitus, lateral view; G. carapace, anterior view; H. sternum, ventral view. Allotype female (PBI_OON 00005598): D. habitus, dorsal view; E. habitus, ventral view; F. habitus, lateral view; I. epigynum, ventral view.

Fig. 24. Ischnothyreus hoplophorus, sp. nov. Holotype male (PBI_OON 00005845): A. left palp, prolateral view; B. left palp, retrolateral view; C. left chelicerae, anterior view; D. left chelicerae, posterior view. Allotype female (PBI_OON 00005598): E. epigynum, ventral view. Scale lines 5 0.1 mm.