Ischnothyreus arcus, Edward & Harvey, 2014

Edward, Karen L. & Harvey, Mark S., 2014, Australian Goblin Spiders Of The Genus Ischnothyreus (Araneae, Oonopidae), Bulletin of the American Museum of Natural History 2014 (389), pp. 1-144 : 133-139

publication ID	https://doi.org/ 10.1206/865.1
persistent identifier	https://treatment.plazi.org/id/E466E016-FF2E-FF36-FCF0-78D43F07FD8B
treatment provided by	Felipe
scientific name	Ischnothyreus arcus
status	sp. nov.

Treatment

Ischnothyreus arcus View in CoL , new species Figures 74–75 View Fig View Fig , map 15

TYPES: AUSTRALIA: Queensland: Male holotype and female allotype from Bulburin State Forest , 500 m, 24.50000 ° S, 151.58330 ° E (25–28 Mar. 1977, V. Davies, R. Raven), deposited in QM (³ holotype: QM S16137 View Materials , PBI _OON 25959; ♀ allotype: QM S16137 View Materials , PBI _OON 25751) GoogleMaps .

ETYMOLOGY: The specific epithet is the Latin arcus , meaning ‘‘bow’’ ( Brown, 1956), and refers to the distinctive sclerotized shape of the female epigynum region, similar to an archer’s bow.

DIAGNOSIS: The male can be distinguished by the presence of both a medial and basal process on the fang, as well as a stout, simple palpal embolic region (fig. 75A–D). The female epigynal region possesses a distinctive horizontal sclerotized processes above a circular epigynal atrium that resembles an archer’s bow (fig. 75F). The curved edge of this overhanging process has a very slight medial triangular extension.

MALE (PBI_OON 25959, figs. 74A–C, G– H, 75A–E). Total length 1.48. CEPHALO- THORAX: Carapace pale orange, broadly oval in dorsal view, pars cephalica strongly elevated in lateral view, anteriorly narrowed to 0.49 times its maximum width or less, anterolateral corners without extension or projections, surface of elevated portion of pars cephalica smooth, sides strongly reticulate; nonmarginal pars cephalica setae light. Clypeus margin unmodified, straight in front view, low, ALE separated from edge of carapace by less than their radius; setae light. Eyes: ALE largest, ALE circular, PME oval, PLE oval; posterior eye row straight from above; ALE touching, ALE-PLE touching.

Sternum as long as wide, pale orange, uniform, setae light, evenly scattered. Chelicerae, endites, and labium pale orange. Chelicerae straight, anterior face unmodified; promargin of chelicerae with two or three larger denticles; fang shape normal, with prominent basal process and medial process; setae light, densest medially; paturon inner margin with scattered setae, inner margin unmodified. Labium elongated hexagon, not fused to sternum, anterior margin slightly indented at middle, same as sternum in sclerotization. Endites anteromedian tip with one strong, toothlike projection, more heavi- ly sclerotized than sternum. ABDOMEN: Ovoid; dorsum soft portions white. Posterior spiracles not connected by groove. Dorsal scutum pale orange, covering more than 3/4 of abdomen, more than 1/2 to most of abdomen width, middle surface smooth, sides smooth. Postepigastric scutum yellow, covering about 1/3 of the abdominal length. Spinneret scutum not visible, specimen bleached. Dorsum, epigastric area, and post- epigastric area setae light. LEGS: Yellow, without color pattern; patella plus tibia I shorter than carapace. Leg spination: femora: I p0-1-1; II p0-1-0; tibiae: I, II p2-2-0; v2-2-0; metatarsi: I, II p1-1-0; v1-1-0. GENITALIA: Palp proximal segments dark red-brown; embolus dark; femur shorter than trochanter, without posteriorly rounded lateral dilation; patella shorter than femur; cymbium dark red-brown; bulb dark red-brown, more than two times as long as cymbium, stout; embolic region stout, simple, without enlarged or complex processes, rounded transparent membrane visible near distal tip, tip split into a number of small processes, one slightly hook shaped (fig. 75A, B, E).

FEMALE (PBI_OON 25751, figs. 74D–F, I, 75F). Total length 1.85. CEPHALOTHORAX: Chelicerae, endites, and labium yellow. ABDO- MEN: Book lung covers ovoid. Dorsal scutum covering 1/2 to 3/4 of abdomen, between 1/4 and 1/2 abdomen width. Postepigastric scutum pale orange, widely hexagonal, only around epigastric furrow. LEGS: Leg spination: femora: I p0-1-1; II p0-1-0; tibiae: I, II p2-2-0; v2- 2-0; metatarsi: I, II p1-1-0; v1-1-0. GENITA- LIA: Ventral view: strong horizontal scleriotization anterior to rounded epigynal atrium, thick, clearly joins posteriorly directed lateral apodemes, shaped like archer’s bow; heavily sclerotized process overhanging epigynal atrium with sinuous edge, slight medial triangular extension; convoluted duct thicker than apodemes (figs. 74I, 75F).

OTHER MATERIAL EXAMINED: AUS- TRALIA: Queensland: 2 km SE of Mount Deongwar , rainforest, 150 m, 27.21667 ° S, 152.28330 ° E, Oct. 14–Dec. 30, 1998 (G. Monteith, D. Cook, QM S54221 View Materials , PBI _OON 22008), 1 ³ GoogleMaps ; Beerwah Forestry Reserve , heathland, 26.85000 ° S, 152.95000 ° E, Nov. 21, 1990 (M. Glover, QM S19529 View Materials , PBI _OON 21980), 1 ³ GoogleMaps ; Beerwah Forestry Reserve , 26.85000 ° S, 152.95000 ° E, Aug. 29, 1990 (M. Glover, QM S19172 View Materials , PBI _OON 22009), 1 ♀ GoogleMaps ; Beerwah Forestry Reserve (C4), heathland, 26.85000 ° S, 152.95000 ° E, June 26, 1991 (M. Glover, QM S63172 View Materials , PBI _OON 22169), 1 ³ GoogleMaps ; Beerwah Forestry Reserve (D4), 26.85000 ° S, 152.95000 ° E, Sept. 19, 1990 (M. Glover, QM S25219 View Materials , PBI _OON 21956), 1 ♀ GoogleMaps ; Beerwah Forestry Reserve (D5), heathland, 26.85000 ° S, 152.95000 ° E, Apr. 24, 1991 (M. Glover, QM S63181, PBI _OON 22182), 2 ♀ GoogleMaps ; Beerwah Forestry Reserve (D5), heathland, 26.85000 ° S, 152.95000 ° E, Apr. 24, 1991 (M. Glover, QM S63196, PBI _OON 22255), 1 ³ GoogleMaps ; Beerwah Forestry Reserve (C4), 26.85000 ° S, 152.95000 ° E, Feb. 27, 1991 (M. Glover, QM S63577 View Materials , PBI _ OON 21991), 1 ♀ GoogleMaps ; Belmont Hills Bushlands , site 1, dry eucalypt woodland, 80 m, 27.51305 ° S, 153.11805 ° E, Feb. 19, 2004 (Queensland Museum Party, QM S79867), 1 ♀ GoogleMaps ; Belmont Hills Bushlands , site 1, dry eucalypt woodland, 27.50784 ° S, 153.11750 ° E, Mar. 1–31, 2004 (Queensland Museum Party, QM S65179 View Materials , PBI _ OON 6431 ), 1 ♀ GoogleMaps ; Belmont Hills Bushlands , site 1, dry eucalypt woodland, 27.50784 ° S, 153.11750 ° E, June 30– July 28, 2003 (S. Wright, E. Volschenk, QM S62263 View Materials , PBI _OON 22185), 1 ³ GoogleMaps ; Belmont Hills Bushlands , site 1, dry eucalypt woodland, 27.50784 ° S, 153.11750 ° E, Mar. 31–Apr. 29, 2004 (Queensland Museum Party, QM S65209 View Materials , PBI _OON 22194), 2 ³, 1 ♀ GoogleMaps ; Belmont Hills Bushlands , site 1, dry eucalypt woodland, 27.50784 ° S, 153.11750 ° E, Jan. 2–29, 2004 (Queensland Museum Party, QM S65180 View Materials , PBI _OON 22195), 1 ³, 2 ♀ GoogleMaps ; Belmont Hills Bushlands , site 1, dry eucalypt woodland, 27.50784 ° S, 153.11750 ° E, Mar. 1– 31, 2004 (Queensland Museum Party, QM S65177 View Materials , PBI _OON 22197), 1 ³ GoogleMaps ; Belmont Hills Bushlands , site 1, dry eucalypt woodland, 27.50784 ° S, 153.11750 ° E, Apr. 16– May 27, 2003 (C. Burwell, S. Wright, E. Volschenk, QM S62479, PBI _OON 22209), 1 ³, 1 ♀ GoogleMaps ; Belmont Hills Bushlands , site 1, dry eucalypt woodland, 27.50784 ° S, 153.11750 ° E, Jan. 30–Mar. 1, 2004 (Queensland Museum Party, QM S65174 View Materials , PBI _OON 22221), 2 ³, 1 ♀ GoogleMaps ; Belmont Hills Bushlands , site 1, dry eucalypt woodland, 27.50784 ° S, 153.11750 ° E, Dec. 1, 2003, to Jan. 2, 2004 (Queensland Museum Party, QM S65211 View Materials , PBI _OON 22223), 2 ³, 2 ♀ GoogleMaps ; Boggomoss No 3 via Taroom , open forest, 25.43333 ° S, 150.01670 ° E, Nov. 11, 1996, to Jan. 12, 1997 (P. Lawless, QM S37179 View Materials , PBI _OON 22126), 5 ³, 2 ♀ GoogleMaps ; Buhot Creek, Burbank , riparian forest, 27.58783 ° S, 153.16980 ° E, Mar. 4, 2004 (C. Burwell, QM S65199, PBI _OON 22190), 5 ♀ GoogleMaps ; Buhot Creek, Burbank , riparian forest, 27.58783 ° S, 153.16980 ° E, Apr. 17–May 27, 2003 (C. Burwell, S. Wright, E. Volschenk, QM S62246, PBI _OON 22414), 1 ♀ GoogleMaps ; Buhot Creek, Burbank , riparian forest, 27.58783 ° S, 153.16980 ° E, Jan. 2–29, 2004 (Queensland Museum Party, QM S65764 View Materials , PBI _OON 22415), 6 ³, 2 ♀ GoogleMaps ; Buhot Creek, Burbank , riparian forest, 27.58783 ° S, 153.16980 ° E, Dec. 12, 2003, to Jan. 1, 2004 (Queensland Museum Party, QM S65762, PBI _OON 22416), 4 ³, 6 ♀ GoogleMaps ; Buhot Creek, Burbank , riparian forest, 27.58783 ° S, 153.16980 ° E, Jan. 30–Mar. 1, 2004 (Queensland Museum Party, QM S65765 View Materials , PBI _OON 22418), 2 ³ GoogleMaps ; Buhot Creek, Burbank , riparian forest, 27.58783 ° S, 153.16980 ° E, July 28–Sept. 2, 2003 (Queensland Museum Party, QM S65761, PBI _OON 22423), 1 ³, 1 ♀ GoogleMaps ; Buhot Creek, Burbank , riparian forest, 27.58783 ° S, 153.16980 ° E, May 27–June 30, 2003 (S. Wright, E. Volschenk, QM S62925, PBI _ OON 22424), 1 ³ GoogleMaps ; Buhot Creek, Burbank , riparian forest, 27.58783 ° S, 153.16980 ° E, Nov. 6, 2003 (Queensland Museum Party, QM S65766, PBI _OON 22426), 1 ♀ GoogleMaps ; Buhot Creek, Burbank , riparian forest, 27.58783 ° S, 153.16980 ° E, Mar. 1–31, 2004 (Queensland Museum Party, QM S79866 View Materials , PBI _ OON 22836), 2 ♀ GoogleMaps ; Buhot Creek, Burbank , riparian forest, leaf litter, 27.58783 ° S, 153.16980 ° E, Feb. 18, 2004 (Queensland Museum Party, QM S65763 View Materials , PBI _OON 25939), 1 ♀ GoogleMaps ; Buhot Creek, Burbank , riparian forest, 27.58783 ° S, 153.16980 ° E, July 28–Sept. 2, 2003 (Queensland Museum Party, QM S62552, PBI _OON 26292), 1 ³ GoogleMaps ; Bulburin Forestry Nursery NW of Bundaberg, rainforest, leaf litter, 24.51666 ° S, 151.48333 ° E, Mar. 1, 1975 (M. Gray, C. Horseman, AM KS6707 , PBI _OON 26284), 1 ³ GoogleMaps ; Bulburin Forestry Nursery NW of Bundaberg, rainforest, leaf litter under rocks, 24.51666 ° S, 151.48333 ° E, Mar. 1, 1975 (M. Gray, C. Horseman, AM KS6708 , PBI _OON 26286), 2 ♀ GoogleMaps ; Bulburin SF, 500 m, 24.50000 ° S, 151.58330 ° E, Mar. 25–28, 1977 (V. Davies, R. Raven, QM S16137 View Materials , PBI _OON 25752), 5 ³, 3 ♀ GoogleMaps ; Bulimba Creek, Carindale , red gum woodland, 27.50150 ° S, 153.10570 ° E, Mar. 1–31, 2004 (Queensland Museum Party, QM S65202 View Materials , PBI _OON 22211), 3 ♀ GoogleMaps ; Bulimba Creek, Carindale , red gum woodland, 27.50150 ° S, 153.10570 ° E, Jan. 2–29, 2004 (Queensland Museum Party, QM S65200 View Materials , PBI _OON 22234), 2 ♀ GoogleMaps ; Bulimba Creek, Carindale , red gum woodland, 27.50150 ° S, 153.10570 ° E, Jan. 30–Mar. 1, 2004 (Queensland Museum Party, QM S65204 View Materials , PBI _ OON 22250), 3 ♀ GoogleMaps ; Bulimba Creek, Carindale , red gum woodland, 27.50150 ° S, 153.10570 ° E, Dec. 1, 2003, to Jan. 2, 2004 (Queensland Museum Party, QM S65203 View Materials , PBI _OON 22252), 1 ³, 1 ♀ GoogleMaps ; Bulimba Creek, Carindale , red gum woodland, 27.50150 ° S, 153.10570 ° E, Mar. 31–Apr. 29, 2004 (Queensland Museum Party, QM S65201, PBI _OON 22476), 7 ♀ GoogleMaps ; Eight Mile Creek (NQ 31), eucalypt woodland, 18.68333 ° S, 144.70500 ° E, Nov. 6, 1991, to July 26, 1992 (R. Raven, P. Lawless, M. Shaw, QM S24689 View Materials , PBI _OON 21751), 1 ³ GoogleMaps ; Eight Mile Creek (NQ 31), eucalypt woodland, 18.68333 ° S, 144.70500 ° E, Nov. 6, 1991, to July 26, 1992 (R. Raven, P. Lawless, M. Shaw, QM S49313 View Materials , PBI _OON 22413), 1 ♀ GoogleMaps ; Fraser I: Orchid Beach, Eliza Ave (FO2), heathland, 24.96667 ° S, 153.30980 ° E, Aug. 20, 1997 (R. Raven, P. Fishburn, P. Lawless, QM S43432, PBI _OON 21744), 1 ³ GoogleMaps ; Fraser Island, Orchid Beach, Eliza Ave , 24.96666 ° S, 153.31638 ° E, Oct. 1, 1996, to Aug. 20, 1997 (R. Raven, P. Fishburn, P. Lawless, QM 72783 , PBI _OON 22000), 1 ³ GoogleMaps ; Gold Creek Reservoir, Brookfield , closed forest, leaf litter, 100 m, 27.50000 ° S, 152.91670 ° E, Dec. 15, 1980 (V. Davies, R. Raven, QM S12997 View Materials , S16481 View Materials – S16483 View Materials , S16485 View Materials ), 2 ³, 4 ♀ GoogleMaps ; Gold Creek Reservoir, site 1, open forest, 27.45883 ° S, 152.87200 ° E, Apr. 20–May 26, 2003 (C. Burwell, QM S65194 View Materials , PBI _OON 22188), 1 ³ GoogleMaps ; Gold Creek Reservoir, site 1, open forest, 27.45883 ° S, 152.87200 ° E, Sept. 1–Oct. 01, 2003 (Queensland Museum Party, QM S62431 View Materials , PBI _OON 22192), 1 ♀ GoogleMaps ; Gold Creek Reservoir, site 1, open forest, 27.45883 ° S, 152.87200 ° E, Mar. 31–Apr. 30, 2004 (Queensland Museum Party, QM S65197 View Materials , PBI _OON 22200), 5 ³, 12 ♀ GoogleMaps ; Gold Creek Reservoir, site 1, open forest, 27.45883 ° S, 152.87200 ° E, July 28–Sept. 1, 2003 (Queensland Museum Party, QM S62292 View Materials , PBI _OON 22201), 2 ♀ GoogleMaps ; Gold Creek Reservoir, site 1, open forest, 27.45883 ° S, 152.87200 ° E, Oct. 30, 2003, Queensland Museum Party, QM S65191 View Materials , PBI _OON 22208), 2 ³, 2 ♀ GoogleMaps ; Gold Creek Reservoir, site 1, open forest, 27.45883 ° S, 152.87200 ° E, Jan. 2–30, 2004 (Queensland Museum Party, QM S65193 View Materials , PBI _OON 22237), 1 ³, 1 ♀ GoogleMaps ; Gold Creek Reservoir, site 1, open forest, 27.45883 ° S, 152.87200 ° E, Jan. 30–Mar. 1, 2004 (Queensland Museum Party, QM S63992, PBI _OON 22253), 1 ³ GoogleMaps ; Gold Creek Reservoir, site 1, open forest, leaf litter, 27.45883 ° S, 152.87200 ° E, Oct. 30, 2003 (Queensland Museum Party, QM S65196, PBI _OON 22254), 1 ³ GoogleMaps ; Gold Creek Reservoir, site 1, open forest, 27.45883 ° S, 152.87200 ° E, Mar. 1–29, 2004 (Queensland Museum Party, QM S65195 View Materials , PBI _OON 22259), 5 ³, 3 ♀ GoogleMaps ; Gold Creek Reservoir, site 1, open forest, 27.45883 ° S, 152.87200 ° E, Oct. 1–30, 2003 (Queensland Museum Party, QM S65198 View Materials , PBI _OON 22472), 1 ³ GoogleMaps ; Illaweena Street, Drewvale , scribblygum open forest, 27.63983 ° S, 153.05780 ° E, Mar. 31–Apr. 29, 2004 (Queensland Museum Party, QM S65210 View Materials , PBI _OON 22244), 3 ³, 1 ♀ GoogleMaps ; Illaweena Street, Drewvale , scribbly gum open forest, 27.63983 ° S, 153.05780 ° E, Mar. 31–Apr. 29, 2004 (Queensland Museum Party, QM S67579 View Materials , PBI _ OON 22245), 1 ³, 3 ♀ GoogleMaps ; Illaweena Street, Drewvale , scribbly gum open forest, 27.63983 ° S, 153.05780 ° E, Apr. 17–May 26, 2003 (C. Burwell, S. Wright, E. Volschenk, QM S67575, PBI _OON 22248), 1 ♀ GoogleMaps ; Karawatha Forest , site 6, dry eucalypt woodland, 27.62217 ° S, 153.08730 ° E, Jan. 2–30, 2004 (Queensland Museum Party, QM S65207, PBI _OON 22207), 1 ³, 5 ♀ GoogleMaps ; Karawatha Forest , site 6, dry eucalypt woodland, 27.62217 ° S, 153.08730 ° E, Mar. 31–Apr. 29, 2004 (Queensland Museum Party, QM S65208 View Materials , PBI _OON 22242), 6 ♀ GoogleMaps ; Karawatha Forest , site 6, dry eucalypt woodland, 27.62217 ° S, 153.08730 ° E, Jan. 30–Mar. 1, 2004 (Queensland Museum Party, QM S65206 View Materials , PBI _OON 22262), 4 ♀ GoogleMaps ; Koy Property, Brigooda , vinescrub, leaf litter, 400 m, 26.26667 ° S, 151.41670 ° E, July 3, 1995 (G. Monteith, QM S57845 View Materials , PBI _OON 22170), 1 ³, 1 ♀ GoogleMaps ; Kroombit Tops (Upper TA 47 Creek.) ; 45 km SSW Calliope , rainforest, 24.40381 ° S, 150.94275 ° E, Dec. 9–19, 1983 (V. Davies, J. Gallon, QM S12976 View Materials , S12978 View Materials ), 3 ³ GoogleMaps ; Kroombit Tops, 45 km SSW. of Calliope , rainforest, leaf litter, 890 m, 24.41666 ° S, 151.01666 ° E, Dec. 10, 1983 (G. Monteith, V. Davies, J. Gallon, G. Thompson, QM S12967 View Materials , PBI _ OON 5572 ), 1 ³ GoogleMaps ; Kroombit Tops , 45 km SSW of Calliope, rainforest, leaf litter, 940 m, 24.41666 ° S, 151.05000 ° E, Dec. 10, 1983 (G. Monteith, V. Davies, J. Gallon, G. Thompson, QM S16096 View Materials , PBI _OON 25889), 1 ³ GoogleMaps ; Kroombit Tops , 45 km SSW of Calliope, rainforest, leaf litter, 940 m, 24.41666 ° S, 151.05000 ° E, Dec. 15, 1983 (G. Monteith, V. Davies, J. Gallon, G. Thompson, QM S16015 View Materials , PBI _OON 25905), 2 ³, 1 ♀ GoogleMaps ; Kroombit Tops , 45 km SSW of Calliope, rainforest, 890 m, 24.41666 ° S, 151.01666 ° E, Dec. 10, 1983 (G. Monteith, V. Davies, J. Gallon, G. Thompson, QM 16043 , PBI _OON 26287), 2 ³ GoogleMaps ; Kroombit Tops, Beauty Spot 98, rainforest, 1000 m, 24.41667 ° S, 151.05000 ° E, Dec. 9–19, 1983 (V. Davies, J. Gallon, QM S12981 View Materials , PBI _OON 25908), 2 ♀ GoogleMaps ; Kroombit Tops, Beauty Spot 98, rainforest, 1000 m, 24.41667 ° S, 151.05000 ° E, Dec. 9–19, 1983 (V. Davies, J. Gallon, QM S16044 View Materials , PBI _OON 25947), 3 ♀ GoogleMaps ; Kroombit Tops, Three Moon Creek , rainforest, 1000 m, 24.36666 ° S, 151.01670 ° E, Dec. 9–19, 1983 (V. Davies, J. Gallon, QM 16012 , PBI _OON 25892), 5 ³ GoogleMaps ; Kroombit Tops, Three Moon Scrub , rainforest, leaf litter, 940 m, 24.41667 ° S, 151.05000 ° E, Sept. 30, 1985 (G. Monteith, QM 16025 ), 1 ³, 1 ♀ GoogleMaps ; Mount Archer (DW 13), open forest, 650 m, 23.33333 ° S, 150.58330 ° E, Sept. 4–Nov. 11, 1991 (D. Wallace, R. Raven, QM 46018 , PBI _OON 21755), 1 ♀ GoogleMaps ; Orchid Beach , Fraser I, eucalypt woodland, 24.96667 ° S, 153.31670 ° E, Mar. 7–Oct. 1, 1996 (R.J. Raven, QM 31281 , PBI _OON 21978), 1 ♀ GoogleMaps ; Proserpine, Lilypool site XY18, closed forest, 30 m, 20.50694 ° S, 148.55888 ° E, Feb. 11–16, 2007 (R. Raven, QM 85252 , PBI _OON 23108), 1 ³ GoogleMaps ; Rochedale SF, rainforest, leaf litter, 27.61667 ° S, 153.15000 ° E, Aug. 30, 1979 (R. Raven, V. Davies, QM 16007 , PBI _ OON 5913 ), 1 ♀ GoogleMaps ; Rose Bay, Bowen. Site 33, vine forest, 20.00000 ° S, 148.26670 ° E, July 27–Dec. 2, 1992 (R. Raven, P. Lawless, E. Lawless, M. Shaw, QM 24954 , PBI _OON 22121), 2 ³ GoogleMaps ; Toomba Homestead site, Rainforest on basalt ridge with Brachychiton , 395 m, 19.96736 ° S, 145.57485 ° E, Feb. 13, 2007 (R. Raven, QM 75374 , PBI _OON 25904), 2 ³ GoogleMaps .