Aetana ternate Huber, 2019

Huber, Bernhard A. & Carvalho, Leonardo S., 2019, Filling the gaps: descriptions of unnamed species included in the latest molecular phylogeny of Pholcidae (Araneae), Zootaxa 4546 (1), pp. 1-96 : 71-74

Aetana ternate Huber sp. n.

Figs 265–266 , 275–280 , 288, 291–293

Aetana Ind 114: Eberle et al. 2018 (molecular data); Huber et al. 2018: fig. 7.

Type material. INDONESIA: ♂ holotype, ZFMK ( Ar 20649), Ternate , Danau (Lake) Tolire (0.830°N, 127.311°E), 180–220 m a.s.l., degraded forest in ravine, 12.xi.2009 (S. Sutono) .

Other material examined. INDONESIA: 2♂ 5♀ 3 juvs, ZFMK (Ar 20650–51), and 1♀ 1 juv. in pure ethanol, ZFMK (Ind207), same data as holotype .

Etymology. The species name is derived from the type locality; noun in apposition.

Diagnosis. Easily distinguished from all known congeners by extremely elongated procursus ( Figs 275 , 277 ) and by extremely elongated female internal genitalia and pore plates ( Fig. 280 ).

Description. Male (holotype). MEASUREMENTS. Total length 2.9, carapace width 1.05. Distance PME-PME 330 µm; diameter PME 100 µm; distance PME-ALE 30 µm; AME absent. Leg 1: 40.2 (9.2 + 0.5 + 9.3 + 17.8 + 3.4), tibia 2: 5.5, tibia 3: 3.7, tibia 4: 5.3; tibia 1 L/d: 93.

COLOR (in ethanol). Carapace ochre yellow with dark brown lateral marginal bands and median band including ocular area; clypeus with pair of dark marks distally; sternum monochromous pale ochre-yellow; legs ochre to light brown, tips of femora and tibiae lighter. Abdomen ochre-gray, with many dark marks dorsally and laterally; ventrally with black median line behind gonopore and dark area in front of spinnerets.

BODY. Habitus as in Fig. 265 . Eye triads on short stalks directed towards lateral ( Fig. 276 ). Thoracic furrow absent (only black median line). Clypeus with pair of apophyses close to distal margin ( Fig. 276 ). Sternum wider than long (0.72/0.50), unmodified.

CHELICERAE. As in Fig. 276 , with pair of long lateral apophyses directed towards lateral and anterior, and pair of small proximal lateral processes.

PALPS. As in Figs 275 and 277 ; coxa barely modified (very indistinct ventral process distally), trochanter proximally very weakly sclerotized, distal part with ventral apophysis, femur with distinctive prolateral process directed towards proximal and curved towards ventral, and two further ventral processes proximally ( Figs 278– 279 ); tibia long, with retrolateral trichobothrium in very distal position; tarsus small; procursus consisting of proximal part and two very long hinged distal parts, one strongly sclerotized, the other proximally widened and with membranous ventral lamina (arrow in Fig. 277 ), distally slender and weakly sclerotized, ending in simple pointed tip; genital bulb very weakly sclerotized (slightly collapsed in all males seen), with short tubular embolus as only process.

LEGS. Without spines; few vertical hairs; with curved hairs on all metatarsi and on tibiae 1 and 2; retrolateral trichobothrium of tibia 1 at 2.5%; tibia 1 without prolateral trichobothrium (present on other tibiae); tarsus 1 with ~30 pseudosegments, distally fairly distinct.

Male (variation). Tibia 1 in 2 other males: 7.8, 8.2.

Female. In general similar to male ( Fig. 266 ) but eye triads closer together (distance PME-PME 170 µm), clypeus unmodified, carapace median mark wider, sternum not monochromous but laterally light brown, legs with dark rings on femora (subdistally) and tibiae (proximally and subdistally), abdomen anteriorly above pedicel with slightly sclerotized area acting against barely modified area on carapace (small median elevation). Tibia 1 in 6 females: 5.3–6.8 (mean 6.2). Epigynum very long ( Figs 288, 291 ), resulting in very posterior position of genital opening; anterior plate barely modified in anterior part, with strong transversal ridges in posterior part; posterior plate simple, apparently without pockets. Internal genitalia with very long pore plates (actually fields of pores fused anteriorly), in posterior part with sclerotized folds and median S-shaped duct, apparently without internal pockets ( Figs 280 , 292–293 ).

Distribution. Known from type locality only ( Fig. 349 ).

Natural history. The spiders were found in a degraded forest in a ravine leading into the lake; their domed webs had a diameter of ~ 30 cm and were built in sheltered spaces among rocks near the ground.

Figures

FIGURE 349. Collecting sites of newly described species of Aetana in eastern Indonesia and of Giloloa sofifi sp. n. on Halmahera. Red mark: two undescribed species deposited in Bernice P. Bishop Museum, Honolulu, Hawaii.

FIGURES 264–271. Giloloa sofifi sp. n. and Aetana spp. live specimens. 264. Giloloa sofifi sp. n., male from near Sofifi. 265–266. Aetana ternate sp. n., male and female from Ternate. 267–269. Aetana mokwam sp. n., male and female from Mokwam. 270–271. Aetana ondawamei sp. n., male and female with egg sac from Gunung Meja.

FIGURES 275–276. Aetana ternate sp. n. (ZFMK, Ar 20650), left male palp, prolateral view, and male prosoma and chelicerae, oblique frontal view. em, embolus; gb, genital bulb; pr, procursus. Scale lines: 0.5.

FIGURES 277–280. Aetana ternate sp. n. (ZFMK, Ar 20650–51). 277. Left male palp, retrolateral view. 278–279. Left male palpal trochanter and femur, retrolateral (slightly dorsal) and ventral views. 280. Cleared female genitalia, dorsal view. em, embolus; fe, femur; gb, genital bulb; pr, procursus; tr, trochanter. Scale lines: 0.5.

FIGURES 288–293. Aetana spp., female abdomens in ventral views, epigynum in ventral view, and cleared female genitalia in ventral and dorsal views. 288, 291–293. A. ternate sp. n. (ZFMK, Ar 20651). 289. A. mokwam sp. n. (ZFMK, Ar 20654). 290. A. ondawamei sp. n. (ZFMK, Ar 20659). Scale lines: 0.5.