DISTICHODONTIDAE Günther, 1864
publication ID |
https://doi.org/ 10.1590/1982-0224-2023-0086 |
publication LSID |
lsid:zoobank.org:pub:3A8C03AC-C66B-49B5-9269-E9A26ABA7EBF |
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https://treatment.plazi.org/id/E118B43D-781C-FED7-FE54-FD5C359CFB20 |
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Felipe |
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DISTICHODONTIDAE Günther, 1864 |
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DISTICHODONTIDAE Günther, 1864 View in CoL View at ENA
T Musschoot
Distichodontina Günther, 1864:280. Type genus: Distichodus Müller, Troschel, 1844 . Additional comments: Géry (1977:58) used the stem Distichod-.
Ichthyborina Günther, 1864:280. Type genus: Ichthyborus Günther, 1864 . Additional comments: Boulenger (1904b:576) changed the stem to Ichthyobor- but it is an unjustified emendation ( Ichthyoborus is preoccupied by Ichthyoborus Kaup, 1845 in birds).
Phagonina Günther, 1865a:175. Type genus: Phago Günther, 1865 . Additional comments: Eigenmann (1909b:255) changed the stem to Phag-.
Nannocharacina Günther, 1867:112. Type genus: Nannocharax Günther, 1867 .
Xenocharacinae Gill, 1893:131. Type genus: Xenocharax Günther, 1867 .
Neolebiinae Eigenmann, 1909b:253. Type genus: Neolebias Steindachner, 1894 .
Hemistichodontinae Regan, 1911:22. Type genus: Hemistichodus Pellegrin, 1900 .
Nannaethiopidi Hoedeman, 1956a:12. Type genus: Nannaethiops Günther, 1872 .
Remarks: Vari (1979:339) considered the Ichthyboridae (or Ichthyborinae) to be included within the Distichodontidae , therefore acting as first reviser in establishing priority.
Belonophago Giltay, 1929:271 . Type species: Belonophago hutsebouti Giltay, 1929 . Type by original designation. Gender: Masculine.
Belonophago hutsebouti Giltay, 1929 Belonophago hutsebouti Giltay, 1929:272 , figure. Type locality: Buta ( Congo Belge). [ Democratic Republic of the Congo]. Holotype: RMCA 19816. Distribution: 534, 535, 537, 540, 546 (Daget, Gosse, 1984; RMCA fish collection; CUMV fish collection). Belonophago tinanti Poll, 1939 Belonophago tinanti Poll, 1939:25 , figs. 10–11. Type locality: Léopoldville and Kunungu (Bolobo), Congo Belge. [ Democratic Republic of the Congo].
Syntypes: RMCA 39438 (1) Kinshasa, 41740–59 (20) Kinshasa, 43896–906 (11) Kinshasa, 44277–308 (30) Kinshasa, 52483–88 (6) Kunungu, 55104 (1) Kinshasa. [Based on the collection number, RMCA 44277–308 should contain 32 syntypes. However, a count of the specimens revealed only 30 are present, with a note mentioning that a single specimen was exchanged with the BMNH. The exchanged specimen agrees to BMNH 1939.3.20.1, but it cannot be determined if this specimen was (still) part of the syntype series as used for the description of B. tinanti , despite it being exchanged as a specimen of B. hutsebouti ].
Distribution: 535, 546, 548 (Poll, 1939; Daget, Gosse, 1984; RMCA fish collection; SMF fish collection).
Congocharax Matthes, 1964b:75 . Type species: Hemigrammocharax olbrechtsi Poll, 1954 . Type by original designation. Gender: Masculine.
Congocharax olbrechtsi (Poll, 1954)
Hemigrammocharax olbrechtsi Poll, 1954:59 View in CoL , fig. 3. Type locality: Yangambi , riv. Lusambila, Congo Belge. [ Democratic Republic of the Congo]. Holotype: RMCA 77451 View Materials .
Distribution: 537 (Poll, 1954; Matthes, 1964b; Daget, Gosse, 1984).
DISTICHODUS Müller, Troschel, 1844
Distichodus Müller, Troschel, 1844:87 . Type species: Characinus nefasch Lacepède, 1803 . Type by monotypy. Gender: Masculine.
Distichodomicrura Fowler, 1936:266 . Type species: Distichodus maculatus Boulenger, 1898 . Type by original designation. Gender: Feminine. Additional comments: Originally proposed as a subgenus of Distichodus .
Distichodura Fowler, 1936:266 . Type species: Distichodus affinis Günther, 1873 . Type by original designation. Gender: Feminine. Additional comments: Originally proposed as a subgenus of Distichodus .
Distichodus affinis Günther, 1873
Distichodus affinis Günther, 1873:144 . Type locality: River Congo, Angola. [Type locality given as “Boma, lower Congo, Congo (Brazzaville)” in the BMNH (now NHMUK) online collection database (although Boma is not located in Congo-Brazzaville but in the Democratic Republic of the Congo) and as “Boma” in Fricke et al. (2022). Boma is located on the right bank of the lower Congo River in Democratic Republic of the Congo, whose lower section (downstream from Matadi and including the section where Boma is located) forms the border between Angola (left bank) and Democratic Republic of the Congo (right bank)]. Holotype: BMNH 1873.7 .28.24.
Distichodus abbreviatus Peters, 1877b:845 View in CoL . Type locality: Congo. Holotype: ZMB 9835 View Materials .
Distribution: 534, 535, 536, 537, 539, 540, 541, 546, 548, 550 (Daget, Gosse, 1984; RMCA fish collection).
Distichodus altus Boulenger, 1899
Distichodus altus Boulenger, 1899b:80 , Pl. 35 (fig. 1). Type locality: Lac Léopold II ( Kutu et Utanda), Congo. [ Democratic Republic of the Congo]. Syntypes: BMNH 1899.9 . 26.113 (1) Ibali; RMCA 733 View Materials to 734 View Materials (1, 1).
Distribution: 520, 534, 535, 537, 546, 547, 548 (Pellegrin, 1904b; Daget, Gosse, 1984; RMCA fish collection; MNHN fish collection; AMNH fish collection).
Distichodus antonii Schilthuis, 1891
Distichodus antonii Schilthuis, 1891:89 View in CoL . Type locality: Bayari-sea , Congo. [ Democratic Republic of the Congo]. Holotype: BMNH 1899.9 .6.10. [See Moelants et al. (2014) for notes on the identification of the holotype].
Distribution: 534, 535, 536, 537, 539, 540, 545, 546, 548, 549, 550 (Moelants et al., 2014).
Distichodus atroventralis Boulenger, 1898
Distichodus atroventralis Boulenger, 1898b:28 , Pl. 13 (fig. 2). Type locality: Boma (2) et Lac Bayari (1), Bas Congo. [Lectotype from: “ DRC: Lake Bayari, Congo River, Lower Congo”. Democratic Republic of the Congo]. Lectotype: BMNH 1891.12 .29.13. [Designated by Moelants et al., 2018:182].
Distribution: 534, 535, 537, 540, 546, 548, 549, 550 (Moelants et al., 2014; Boulenger, 1909a; Gosse, 1968).
Remarks: Redescription of species presented by Moelants et al., 2018.
Distichodus brevipinnis Günther, 1864
Distichodus brevipinnis Günther, 1864:361 View in CoL . Type locality: Chartoum , Upper Nile. [ Sudan]. Syntypes: BMNH 1862.6.17.104–106 (3).
Distribution: 505, 506, 507, 508, 509, 516, 520, 522 (Gosse, Coenen, 1990; Gosse et al., 2003).
Distichodus decemmaculatus Pellegrin, 1926
Distichodus decemmaculatus Pellegrin, 1926:160 View in CoL . Type locality: Kunungu (bassin du Congo), Congo Belge. [ Democratic Republic of the Congo]. Holotype: RMCA 19697 View Materials .
Distribution: 535, 537, 540, 546 (David, Poll, 1937; Lambert, 1961; Stiassny et al., 2021a).
Distichodus engycephalus Günther, 1864
Distichodus engycephalus Günther, 1864:361 View in CoL . Type locality: Chartoum , Upper Nile. [ Sudan]. Syntypes: BMNH 1862.6.17.80–81 (2).
Distichodus touteei Pellegrin, 1906d:472 View in CoL . Type locality: Niger View in CoL : entre Gaya, Boussa et Badjibo. [ Niger View in CoL River between Gaya ( Niger View in CoL ), Boussa (= Bussa, Nigeria) and Badjibo (upstream from Jebba, Nigeria)]. Syntypes: MNHN 1900-0103 to 0105 (3).
Distribution: 505, 506, 507, 508, 516, 517, 518, 520, 522, 523 (Gosse et al., 2003; Vari , 2007; Boulenger, 1907c).
Distichodus fasciolatus Boulenger, 1898
Distichodus fasciolatus Boulenger, 1898b:28 , Pl. 14. Type locality: Boma GoogleMaps , Manyanga, Léopoldville (bassin du Congo). [Lectotype from: “Boma ( DRC) (± 5°50’S, 13°03’E)”. Democratic Republic of the Congo]. Lectotype: RMCA 23 View Materials . [Designated by Moelants et al. (2014)].
Distribution: 534, 535, 536, 537, 538, 540, 541, 545, 546, 548, 549, 550 (Kisekelwa et al., 2020; Moelants et al., 2014).
Distichodus hypostomatus Pellegrin, 1900
Distichodus hypostomatus Pellegrin, 1900b:351 View in CoL . Type locality: Adouma (Ogôoué), Congo français. [ Gabon]. Syntypes: MNHN 1886-0369 View Materials to 0371 View Materials (3).
Distribution: 532, 533 ( Vari , 2007).
Remarks: Distichodus hypostomatus Vaillant in Rivière 1886 is an unavailable name.
Distichodus ingae Moelants, Snoeks, 2018
Distichodus ingae Moelants, Snoeks , in Moelants GoogleMaps , Snoeks GoogleMaps , Vreven GoogleMaps , 2018:189, figs. 5a,b, 11. Type locality: DRC, Congo River GoogleMaps near Inga, a few km upstream and on the opposite side of river from Inga hydroelectric dam, 5°27’30”S, 13°36’00”E. [ Democratic Republic of Congo]. Holotype: MCZ 50439 View Materials .
Distribution: 549 (Moelants et al., 2018).
Distichodus kasaiensis Vreven, Moelants, Snoeks, 2018
Distichodus kasaiensis Vreven, Moelants, Snoeks , in Moelants GoogleMaps , Snoeks GoogleMaps , Vreven GoogleMaps , 2018:187, figs. 9–10. Type locality: DRC: Lukoshi River GoogleMaps above and below rapids of Mukanda GoogleMaps or Kamakamaka, Lulua subbasin, approximately 14.5 km in a straight line SSW of Kasaji, 10°30’44”S, 23°23’08”E. [Democratic Republic of Congo]. Holotype: RMCA B2-030 View Materials - P-1389 .
Distribution: 546 (Moelants et al., 2018).
Distichodus kolleri Holly, 1926
Distichodus kolleri Holly, 1926:158 . Type locality: Sanaga und Bam-fluss, einem Nebenflusse des Sanaga in Kamerun. [“Sanaga and Bam River, an affluent of the Sanaga in Cameroon ”]. Syntypes: NMW 7167 View Materials (1), 7168–72 (5).
Distichodus vexillifer Pellegrin, 1927a:139 . Type locality: Noun, près de Bafousam (Cameroun) et Nanga-Tboko (Moyenne Sanaga-Cameroun). [Noun, near Bafousam and Nanga-Tboko (middle Sanaga River) in Cameroon]. Syntypes: MNHN 1927-0087 View Materials to 0089 View Materials (1, 1, 1) Noun River , 1927-0090 (1) Sanaga River. [Based on the online collection database of the MNHN, MNHN 1927-0087 View Materials to 0089 View Materials were collected at Bafousan (Sanaga River basin, Cameroon) and MNHN 1927-0090 View Materials was collected at Nanga-Eboko (Sanaga River basin, Cameroon)].
Distribution: 505 ( Vari , 2007).
Distichodus langi Nichols, Griscom, 1917
Distichodus langi Nichols, Griscom, 1917:687 View in CoL , Pl. 69. Type locality: Faradje , Belgian Congo. [ Democratic Republic of the Congo]. Holotype: AMNH 5915 View Materials .
Distribution: 536, 537, 540, 541 (Kisekelwa et al., 2020; Abwe et al., 2019; Moelants et al., 2014).
Distichodus lusosso Schilthuis, 1891
Distichodus lusosso Schilthuis, 1891:90 . Type locality: Bayari-sea. [The type locality given with the original description is “Bayari-Sea”, while the introductory part of the publication mentioned that the specimens reported upon originated from “Boma, lower Congo, and for the greater part from Kinshassa, Stanley Pool, upper Congo, near Brazzaville”. Democratic Republic of Congo]. Syntypes: BMNH 1891.12.29.13 (1), 1899.9.6.11 (1). [(1) BMNH 1898.11.17.18–19 (2), listed as syntypes in Daget, Gosse (1984) and Eschmeyer (1998), are not type specimens. They are much larger than those mentioned in the original description of the species (240–360 mm TL versus at most 168 mm TL, respectively) and are from another collector (“Pres? by the Secretary of State, Congo Free State ”, versus collected by Mr. A Greshoff). In addition , the larger specimens originated from “Manyanga” and were originally registered in the BMNH collection as D. leptorhynchus Boulenger, 1897 . (2) BMNH 1891.12.29.13, which is here identified as the smaller of the two syntypes mentioned in the original description of D. lusosso , measuring about 63.5 mm SL, is also syntype of D. atroventralis Boulenger 1898 . BMNH 1899.9.6.11, although slightly smaller than the size given in the original description (about 152 mm TL instead of 168 mm TL), is believed to be the larger of the two syntypes. Both BMNH 1891.12.29.13 and BMNH 1899.9.6.11 are catalogued in the BMNH register as types, the former from “lower Congo”, the latter from “Kinshassa, Stanley Pool”].
Distichodus leptorhynchus Boulenger, 1897b:425 View in CoL . Type locality: Stanley Falls , upper Congo. [ Democratic Republic of the Congo]. Holotype: BMNH 1897.9 .30.17.
Distribution: 534, 535, 536, 537, 539, 540, 541, 545, 546, 548, 549 (Daget, Gosse, 1984; Ibala Zamba, 2010; RMCA fish collection).
Remarks: Vernacular name: Lusosso . Distichodus fasciatus Vaillant in Rivière 1886 is an unavailable name.
Distichodus maculatus Boulenger, 1898
Distichodus maculatus Boulenger, 1898b:27 View in CoL , Pl. 13 (fig. 1). Type locality: Manyanga , Congo. [ Democratic Republic of the Congo]. Holotype: RMCA 86 View Materials .
Distribution: 534, 535, 536, 537, 540, 542, 543, 544, 545, 546, 548, 549 (Daget, Gosse, 1984; RMCA fish collection; Eccles, 1992).
Distichodus mbiniensis Schmidt, Knobloch, Barrientos, 2021
Distichodus mbiniensis Schmidt, Knobloch, Barrientos, 2021:305 , fig. 7. Type locality: Equatorial Guinea, Wele-Nzas, Rio Ntoro at Aconibe Acurenam Rd. GoogleMaps , 668 m elev, 1.29091°N, 10.9174°E. Holotype: USNM 451317 View Materials .
Distribution: 533 (Schmidt et al., 2021).
Distichodus microps Schmidt, Knobloch, Barrientos, 2021
Distichodus microps Schmidt, Knobloch, Barrientos, 2021:301 , fig. 5. Type locality: Equatorial Guinea, Litoral, Rio Mbia at Menzong GoogleMaps , 32 m elev, 2.07529°N, 9.92378°E. Holotype: USNM 451325 View Materials .
Distribution: 533 (Schmidt et al., 2021).
Distichodus mossambicus Peters, 1852
Distichodus mossambicus Peters, 1852:275 View in CoL . Type locality: Mossambique. [ Daget , Gosse (1984) specify the type locality as “ Tette, Zambezi River ”. Mozambique]. Syntypes: ZMB 3564 View Materials (1), 6613 (1).
Distichodus macrolepis Günther, 1864:362 View in CoL . Type locality: Shire River , near the island of Dakanamois. [ Malawi]. Holotype: BMNH 1864.1 .9.62 (skin).
Distribution: 558, 561, 576 (Skelton, 2001; Moelants et al., 2014).
Distichodus nefasch (Bonnaterre, 1788)
Salmo nefasch Bonnaterre, 1788:169 . Type locality: Nile. [Type locality is probably situated in Egypt: Bonnaterre (1788) mentions the specific name “nefasch ” is used in reference to the common name for the species as used by the Egyptians]. No types known.
Salmo aegyptius Gmelin, 1789:1386 . Type locality: Aegypto. [ Egypt]. No types known.
Characinus nefasch Lacepède, 1803:270 . Type locality: Nile River. No types known. Bertin (1948) incorrectly attributed type status to the following specimens: MNHN A-8603 and 9878 (2, dry), MNHN A-9587 (1), A-9588 (4), A-7929 (1), A-9879 to A-9881 (1, 1, 1, 1), A-8602 (1).
Distichodus rudolphi Günther, 1896a:223 View in CoL . Type locality: Lake Rudolf [Lake Turkana]. Syntypes: ANSP 14559 View Materials (1), BMNH 1895.12 .31.71 (1).
Distribution: 522, 523, 530 (Boulenger, 1907c; Lévêque et al., 1991; Daget, Gosse, 1984).
Remarks: Salmo niloticus Hasselquist, 1762 is an unavailable name.
Distichodus noboli Boulenger, 1899
Distichodus noboli Boulenger, 1899b:81 View in CoL , Pl. 35 (fig. 2). Type locality: Kutu (Lac Léopold II), Congo. [ Democratic Republic of the Congo]. Holotype: RMCA 735 View Materials .
Distribution: 534, 535, 537, 538, 546, 547, 548 (Daget, Gosse, 1984; RMCA fish collection; CUMV fish collection; SMF fish collection; Boulenger, 1909a; Matthes, 1964b).
Distichodus notospilus Günther, 1867
Distichodus notospilus Günther, 1867:114 View in CoL [6]. Type locality: Gaboon [ Gabon]. Syntypes: BMNH 2013.4.17.2–4 (3).
Distribution: 505, 532, 533, 534, 537, 546, 550 ( Vari , 2007; RMCA fish collection; Daget, Gosse, 1984; Schmidt et al., 2021:309).
Remarks: Geographic variation and biogeography discussed by Schmidt et al., 2021.
Distichodus petersii Pfeffer, 1896
Distichodus petersii Pfeffer, 1896:41 View in CoL . Type locality: Kingani-Fluss , Deutsch Ostafrika. [“Kingani River, German East Africa”. Tanzania]. Syntypes: BMNH 1908.1 . 27.3 (1), ZMH H473 View Materials , ZMB 13677 View Materials (4).
Distichodus albini Steindachner, 1914b:537 View in CoL . Type locality: Deutsch-Ostafrika. [ Steindachner (1916) provided a more elaborate description of the species, including details on the type locality: “ Ulanga-Fluss bei Ifakara , Deutsch-Ostafrika ” (Ulanga River near Ifakara, German East Africa = Tanzania)]. Holotype: NMW 68682 View Materials .
Distribution: 564 (Daget, Gosse, 1984).
Distichodus polli Abwe, Snoeks, Chocha Manda, Vreven, 2019
Distichodus polli Abwe, Snoeks, Chocha Manda, Vreven, 2019:86 , figs. 5, 6a. Type locality: Democratic Republic of the Congo: Kasaï-Oriental Province GoogleMaps , Kasai GoogleMaps basin: Bushimaie River, Lula, approximately 7°20’ S 23°08’ E. Holotype: RMCA 153493 View Materials .
Distribution: 545, 546 (Abwe et al., 2019).
Distichodus rostratus Günther, 1864
Distichodus rostratus Günther, 1864:360 View in CoL . Type locality: Lower Nile River and West Africa. Syntypes: BMNH uncat. (stuffed, apparently lost) West Africa , BMNH 1861.9 .9.25 (1) Lower Nile .
Distichodus martini Steindachner, 1870:549 View in CoL [17], Pl. 3 (figs. 2–2a). Type locality: Dagana und Podor, Senegal. Syntypes: NMW 68683 View Materials (9), 77251 (6).
Distribution: 505, 506, 507, 508, 509, 514, 515, 516, 517, 518, 520, 521, 522, 523 (Gosse et al., 2003; Vari , 2007; Boulenger, 1907c).
Distichodus rufigiensis Norman, 1922
Distichodus rufigiensis Norman, 1922:686 View in CoL . Type locality: Mpanganye , Rufigi River, Tanganyika Territory. [ Tanzania]. Syntypes: BMNH 1922.4.18.15–17 (3).
Distribution: 564 (Daget, Gosse, 1984).
Distichodus schenga Peters, 1852
Distichodus schenga Peters, 1852:276 View in CoL . Type locality: Mossambique. [ Daget , Gosse (1984) specify the type locality as “ Tette, Zambezi River,” Mozambique]. Syntypes: ZMB 3565 View Materials (now 3).
Distribution: 558, 561, 576 (Skelton, 2001).
Distichodus sexfasciatus Boulenger, 1897
Distichodus sexfasciatus Boulenger, 1897b:425 View in CoL . Type locality: Stanley Falls , upper Congo. [ Democratic Republic of the Congo]. Holotype: BMNH 1897.9 .30.16.
Distichodus tanganicae Worthington, Ricardo, 1937:1083 View in CoL , fig. 2. Type locality: Lake Tanganyika. [The lack of details on the type locality is explained in the original description, stating that “it is extremely unfortunate, however, that Christy’s note-books have been mislaid, so that there are practically no data as the where, when, or how the fish were caught”]. Holotype: BMNH 1936.6 .15.411.
Distribution: 534, 535, 536, 537, 539, 540, 541, 542, 545, 546, 547, 548, 549, 550 (RMCA fish collection; Boulenger, 1909a; Poll, 1939; Poll, 1953; Daget, Stauch, 1963; Poll, 1967; Gosse, 1968; Poll, 1976; Roberts, Stewart, 1976; Banister, Bailey, 1979; Vivien, 1991; Monsembula Iyaba, Stiassny, 2013; Kisekelwa et al., 2020).
Distichodus teugelsi Mamonekene, Vreven, 2008
Distichodus teugelsi Mamonekene, Vreven, 2008:99 , fig. 1. Type locality: Republic of Congo: Congo River GoogleMaps basin: Léfini River GoogleMaps , 3km upstream of confluence with the Louna River, Réserve Naturelle Lésio-Louna, 3°00’S, 15°30’E. Holotype: RMCA 2004-46 View Materials - P-0005 .
Distribution: 534, 537 (Mamonekene, Vreven, 2008).
Dundocharax Poll, 1967:129 . Type species: Dundocharax bidentatus Poll, 1967 . Type by original designation. Gender: Masculine.
Dundocharax bidentatus Poll, 1967
Dundocharax bidentatus Poll, 1967:129 View in CoL , fig. 54. Type locality: Riv. Lucoge, Angola. Holotype: MD 6500 .
Distribution: 546 (Poll, 1967; Daget, Gosse, 1984).
EUGNATHICHTHYS Boulenger, 1898
Eugnathichthys Boulenger, 1898b:25 . Type species: Eugnathichthys eetveldii Boulenger, 1898 . Type by original designation. Gender: Masculine.
Eugnathichthys eetveldii Boulenger, 1898 Eugnathichthys eetveldii Boulenger, 1898b:26 , Pl. 12. Type locality: Léopoldville,
Boma, Nouvelle-Anvers et Upoto, Congo. [ Democratic Republic of the Congo]. Syntypes: BMNH 1898.11.17.8–9 (2) Leopoldville; RMCA 26 (1) Boma, 121 (1) Leopoldville, 148 (1) Upoto.
Eugnathichthys intermedius Pappenheim , in Pappenheim , Boulenger , 1914:234, Pl. 4 (fig. 5). Type locality: Aruwimi bei Basoko, Congo. [ Democratic Republic of the Congo]. Holotype: ZMB 19043 View Materials .
Distribution: 534, 535, 536, 537, 538, 539, 540, 545, 546, 548, 549, 550 (RMCA fish collection; AMNH fish collection; Boulenger, 1898b; Poll, 1939; Daget, Stauch, 1963; Gosse, 1968; Poll, 1976; Banister, Bailey, 1979; Stiassny et al., 2013; Nichols, Griscom, 1917).
Eugnathichthys macroterolepis Boulenger, 1899
Eugnathichthys macroterolepis Boulenger, 1899b:75 , Pl. 34 (fig. 1). Type locality: Chiloango. [ The BMNH online collection database specifies the type locality as the “ Loange River , Belgian Congo”. The Loange River is a tributary of the Chiloango in Democratic Republic of the Congo]. Holotype: BMNH 1899.6 .28.16.
Distribution: 532, 534, 535, 536, 537, 538, 539, 540, 541, 545, 546, 548, 549, 550 (RMCA fish collection; Boulenger, 1909a; Poll, 1976; Matthes, 1964b; David, Poll, 1937; Nichols, Griscom, 1917; Daget, Stauch, 1963; Gosse, 1968; Vivien, 1991; Vari , 2007; Stiassny et al., 2013; Kisekelwa et al., 2020).
Eugnathichthys virgatus Stiassny, Denton, Monsembula Iyaba, 2013
Eugnathichthys virgatus Stiassny, Denton, Monsembula Iyaba, 2013:483 , figs. 3–5A. Type locality: Democratic Republic of Congo, Equateur Province GoogleMaps , Luilaka River at confluence of small stream flowing out of Salonga National Park, 2.6713°S, 21.71452°E. Holotype: AMNH 241648 View Materials .
Distribution: 534, 537 (Stiassny et al., 2013).
Ichthyborus Günther, 1864:362 . Type species: Ichthyborus microlepis Günther, 1864 . Type by subsequent designation by Jordan (1919b). Gender : Masculine.
Psalidostoma Kner, 1864:99 . Type species: Psalidostoma caudimaculatum Kner, 1864 View in CoL . Type by monotypy. Gender: Neuter. Additional comments: Preoccupied by Psalidostoma Minding, 1832 in fishes, replaced by Onouphrios Whitley, 1951a .
Neoborus Boulenger, 1899b:78 . Type species: Neoborus ornatus Boulenger, 1899 View in CoL . Type by monotypy. Gender: Masculine. Additional comments: Preoccupied by Neoborus Distant, 1884 in Hemiptera , replaced by Phagoborus Myers, 1924b .
Phagoborus Myers, 1924b:397 . Type species: Neoborus ornatus Boulenger, 1899 . Type by being a replacement name. Gender: Masculine. Additional comments: Replacement for Neoborus Boulenger, 1899 preoccupied by Neoborus Distant, 1884 in Hemiptera .
Gavialocharax Pellegrin, 1927b:390 . Type species: Gavialocharax monodi Pellegrin, 1927 . Type by monotypy. Gender: Masculine.
Ra Whitley, 1931:334. Type species: Ichthyborus microlepis Günther, 1864 . Type by being a replacement name. Gender: Feminine. Additional comments: Unneeded replacement for Ichthyborus Günther, 1864 , not preoccupied (Daget, Gosse, 1984:197).
Onouphrios Whitley, 1951a:68 . Type species: Psalidostoma caudimaculatum Kner, 1865 View in CoL . Type by being a replacement name. Gender: Masculine. Additional comments: Replacement for Psalidostoma Kner, 1864 preoccupied by Psalidostoma Minding, 1832 in fishes.
Ichthyborus besse (Joannis, 1835) View in CoL
Characinus besse Joannis, 1835 :no page number, Pl. 10. Type locality: Nil , Egypte [Nile, Egypt]. Syntypes: MNHN A-9721 (2, poor condition).
Ichthyborus microlepis Günther, 1864:363 View in CoL . Type locality: Chartoum , Upper Nile. [ Sudan]. Syntypes: BMNH 1863.3.9.22–23 (2).
Psalidostoma caudimaculatum Kner, 1864:100 View in CoL , Pl. Type locality: Weissen Nil. [White Nile River]. Holotype: lost.
Ichthyoborus besse congolensis Giltay, 1930:395 , figs. 2–3. Type locality: Nyonga (Haut-Luapula, Congo Belge). [ Democratic Republic of the Congo]. Holotype: RMCA 20555 View Materials .
Distribution: 505, 520, 522, 523, 524, 540 (Boulenger, 1907c; Gosse et al., 2003; Decru et al., 2017).
Ichthyborus monodi (Pellegrin, 1927)
Gavialocharax monodi Pellegrin, 1927b:391 View in CoL . Type locality: Yabassi , Cameroun. [ Cameroon]. Syntypes: MNHN 1926-0289 View Materials to 0291 View Materials (1, 1, 1).
Distribution: 506, 517, 518 (Gosse et al., 2003; Vari , 2007).
Ichthyborus ornatus (Boulenger, 1899)
Neoborus ornatus Boulenger, 1899b:78 , Pl. 34 (fig. 4). Type locality: Bikoro (lac Tumba ) et Kutu (lac Léopold II), Congo. [ Democratic Republic of the Congo]. Syntypes: BMNH 1899.9 . 26.85 (1) Kutu , RMCA 712 View Materials to 715 View Materials (1, 1, 1, 1).
Distribution: 534, 535, 537, 538, 540, 546, 547, 548 (RMCA fish collection; CUMV fish collection; Boulenger, 1899b; Pellegrin, 1928b; David, Poll, 1937; Poll, 1959; Daget, Stauch, 1963; Gosse, 1968; Decru et al., 2017).
Remarks: Ichthyoborus taeniatus Vaillant in Rivière, 1886 is an unavailable name.
Ichthyborus quadrilineatus (Pellegrin, 1904)
Neoborus quadrilineatus Pellegrin, 1904a:219 View in CoL . Type locality: Casamance, à la hauteur de Kolibantan, Guinée portugaise. [ Guinea-Bissau]. Holotype: MNHN 1904- 0054 View Materials .
Distribution: 509, 511 (Gosse et al., 2003).
Mesoborus Pellegrin, 1900a:178 . Type species: Mesoborus crocodilus Pellegrin, 1900 . Type by monotypy. Gender: Masculine.
Champsoborus Boulenger, 1909d:187 . Type species: Champsoborus pellegrini Boulenger, 1909 . Type by monotypy. Gender: Masculine.
Mesoborus crocodilus Pellegrin, 1900
Mesoborus crocodilus Pellegrin, 1900a:179 View in CoL . Type locality: Diélé sur l’Alima , Congo français. [ Republic of the Congo]. Holotype: MNHN 1886-0377 View Materials .
Champsoborus pellegrini Boulenger, 1909d:187 View in CoL . Type locality: Bumba River at Assobam , South Cameroon; Ubanghi River. [No information is provided on the exact origin of the Ubanghi River specimen in the original description, but the RMCA fish collection specifies that it comes from “Banzyville, river Ubangi, Etat Indépendent du Congo”. Democratic Republic of the Congo]. Syntypes: BMNH 1909.7.9.47–48 (2), RMCA 1301 View Materials (1).
Distribution: 534, 535, 536, 537, 539, 540, 541, 546, 548 (RMCA fish collection; Vivien, 1991; Monsembula Iyaba, Stiassny, 2013; Kisekelwa et al., 2020).
MICROSTOMATICHTHYOBORUS Nichols, Griscom, 1917
Microstomatichthyoborus Nichols, Griscom, 1917:685 . Type species: Microstomatichthyoborus bashforddeani Nichols, Griscom, 1917 . Type by monotypy. Gender: Masculine.
Tristichodus Boulenger, 1920b:19 . Type species: Tristichodus christyi Boulenger, 1920 . Type by monotypy. Gender: Masculine.
Microstomatichthyoborus bashforddeani Nichols, Griscom, 1917
Microstomatichthyoborus bashforddeani Nichols, Griscom, 1917:685 View in CoL , fig. 6. Type locality: Poko , Belgian Congo. [ Democratic Republic of the Congo]. Holotype: AMNH 6329 View Materials .
Tristichodus christyi Boulenger, 1920b:20 View in CoL , fig. 8. Type locality: Poko , Congo belge. [ Democratic Republic of the Congo]. Syntypes: BMNH 1919.9 . 10.141 (1), RMCA 7160 View Materials (1).
Distribution: 534, 535, 536, 537, 540, 546 (RMCA fish collection; CUMV fish collection; Monsembula Iyaba, Stiassny, 2013).
Microstomatichthyoborus katangae David, Poll, 1937
Microstomatichthyoborus katangae David , Poll, 1937:212, fig. 10. Type locality: Nionga, Congo Belge. [ Democratic Republic of the Congo]. Holotype: RMCA 23159 View Materials . [In the original description, a total of 213 specimens are cited: 20 (“including the type”) from Nionga, 171 (“ paratypes ”) from Nionga, 18 from Kabelwe (agrees with RMCA 27054–27069 View Materials (16) and 27822–27823 (2)) and 5 from Kabalo (agrees with RMCA 23211-2315 View Materials ). Following the ICZN, 1999: Art. 73.1, the use of “the type” is a holotype designation. In addition , the use of “ paratypes ” for the other type specimens is an additional argument to believe that the authors wanted to designate a holotype. A closer look at the specimens in the RMCA collection revealed that RMCA 23159–168 View Materials (10 specimens from Nyonga, labelled as syntypes in the RMCA database) contained a tube with a single specimen, with a note saying “type (expl. figure).” Although this specimen is smaller than the 54mm given below the figure in the original description, as are all specimens in this lot, its coloration pattern matches the photograph, and it is to be treated as the holotype. 189 remaining paratype specimens from Nyonga were counted (which is lower than the expected number of specimens based on the collection numbers), including the remaining “ syntypes ” from RMCA 23159–168 View Materials (9), another lot labelled as “ syntypes ” ( RMCA 23169–178 View Materials ;10) and 20 other lots, while the original publication gives 190. All these Nyonga specimens are to be treated as paratypes].
Distribution: 540, 545 (RMCA fish collection; David, Poll, 1937).
Monostichodus Vaillant , in Rivière, 1886:17. Type species: Monostichodus elongatus Vaillant, 1886 . Type by monotypy. Gender: Masculine.
Hemistichodus Pellegrin, 1900b:352 . Type species: Hemistichodus vaillanti Pellegrin, 1900 . Type by monotypy. Gender: Masculine.
Remarks: Taxonomic status of Monostichodus and Hemistichodus is discussed by Musschoot, Snoeks, 2016.
Monostichodus elongatus Vaillant, 1886
Monostichodus elongatus Vaillant , in Rivière, 1886:17. Type locality: Not specified. [Type locality not specified, but all fishes in the original description originated from either the Ogôoué River (Franceville or Adoumas) or the Congo River basin (Mokaka on the Sangua River, Alima River, Nganchou or Brazzaville). No types known.
Hemistichodus vaillanti Pellegrin, 1900b:352 View in CoL . Type locality: Adouma (Ogôoué), Congo français. [Ogooué River at Adouma, Gabon]. Holotype: MNHN 1886- 0376 View Materials .
Distribution: 532 ( Vari , 2007).
Remarks: Taxonomic status is discussed by Musschoot, Snoeks (2016).
Monostichodus lootensi (Poll, Daget, 1968)
Hemistichodus lootensi Poll, Daget, 1968:1060 View in CoL . Type locality: Nkombo , Coquilhatville, Congo central. [ Democratic Republic of the Congo]. Holotype: RMCA 78571 View Materials .
Distribution: 537, 538, 546, 548 (RMCA fish collection; Poll, Daget, 1968; Stiassny et al., 2021a).
Monostichodus mesmaekersi (Poll, 1959)
Hemistichodus mesmaekersi Poll, 1959:85 View in CoL , Pl. 16 (fig. 2A). Type locality: Riv. noire, chenal du Stanley Pool, Congo Belge. [ Democratic Republic of the Congo]. Holotype: RMCA 118973 View Materials .
Distribution: 534, 537, 538, 540, 546 (RMCA fish collection; CUMV fish collection; Poll, 1959; Decru et al., 2017; Stiassny et al., 2021a).
Nannaethiops Günther, 1872a:669 . Type species: Nannaethiops unitaeniatus Günther, 1872 . Type by monotypy. Gender: Masculine.
Nannaethiops bleheri Géry, Zarske, 2003 Nannaethiops bleheri Géry, Zarske, 2003:39 , figs. 1–2. Type locality: Tata Lake in einem Abfluss des Baro River (abfliessend in den Sawba River) nahe Gési, westlich von Aganga, Gambela district, südwestliches Äthiopien, stromgebiet des oberen Weissen Nil nahe der Grenze zum Sudan. [ Ethiopia]. Holotype: MTD F 24823 View Materials . Distribution: 522 (Géry, Zarske, 2003).
Nannaethiops unitaeniatus Günther, 1872
Nannaethiops unitaeniatus Günther, 1872a:670 View in CoL , Pl. 65 (fig. C). Type locality: Gaboon [ Gabon]. Lectotype: BMNH 1872.1 .27.8. [Designated by Poll, Gosse (1963a)].
Distribution: 505, 506, 518, 520, 532, 533, 534, 550 (Poll, Gosse, 1963a; Gosse et al., 2003; Vari , 2007; RMCA fish collection).
Nannocharax Günther, 1867:112 . Type species: Nannocharax fasciatus Günther, 1867 . Type by monotypy. Gender: Masculine.
Hemigrammocharax Pellegrin, 1923:115 (footnote). Type species: Nannocharax ocellicauda Boulenger, 1907 . Type by monotypy. Gender: Masculine. Additional comments: Originally proposed as a subgenus of Nannocharax .
Microdistichodus Pellegrin, 1926:161 . Type species: Microdistichodus uniocellatus Pellegrin, 1926 . Type by monotypy. Gender: Masculine.
Thrissocharax Myers, 1926d:174 . Type species: Nannocharax ocellicauda Boulenger, 1907 . Type by original designation. Gender: Masculine.
Hemigrammonannocharax Holly, 1930:198 , 233. Type species: Nannocharax ocellicauda Boulenger, 1907 . Type by original designation. Gender: Masculine.
Distichodina Fowler, 1935:260 . Type species: Distichodus stigmaturus Fowler, 1935 . Type by original designation. Gender: Feminine. Additional comments: Originally proposed as a subgenus of Distichodus .
Lepinannocharax Fowler, 1936:271 . Type species: Nannocharax pteron Fowler, 1936 . Type by original designation. Gender: Masculine. Additional comments: Originally proposed as a subgenus of Nannocharax .
Nannocharax altus Pellegrin, 1930
Nannocharax altus Pellegrin, 1930:203 . Type locality: Kouilou , Afrique équatoriale française. [Based on the introductory part of the original description, the species was probably collected in the upper Kouilou-Niari basin (including the rivers Louessé, Loula, Lali and Bouenza) in Republic of Congo (= Congo-Brazzaville)]. Syntypes: MNHN 1929-0242 View Materials (17, now 16); RMCA 20646 View Materials (1) ;? MSNM 2018 (1), USNM 92963 View Materials (1).
Distribution: 532 ( Vari , 2007).
Nannocharax angolensis (Poll, 1967)
Hemigrammocharax angolensis Poll, 1967:121 View in CoL , fig. 51. Type locality: Poste de Cuílo GoogleMaps , Xá-Ua, riv. Luíta, 8°02’S, 19°25’E, Angola. Holotype: MD 6222 .
Distribution: 546 (RMCA fish collection; Poll, 1967).
Nannocharax ansorgii Boulenger, 1911
Nannocharax ansorgii Boulenger, 1911a:374 . Type locality: Geba and Culufi Rivers, Portuguese Guinea. [ Guinea-Bissau]. Syntypes: BMNH 1912.4.1.78–82 (5), 1912.4.1.83–87 (5);? MCZ 31505 View Materials (1) ; RMCA 7095 View Materials (1) ; USNM 92960 View Materials (1) ; ZMB 18445 View Materials (6) Culufi River . [Although Fricke et al. (2022) mentioned that USNM 92960 View Materials (labelled as cotype in the USNM online collection database) was formerly a specimen registered under BMNH 1912.4.11.373-382, the latter lot (or the remaining specimens from this lot) could not be traced in the Natural History Museum in London (J. Maclaine, pers. comm.)] .
Hemigrammocharax polli Roman, 1966:81 , Pl. 6 (fig. 12). Type locality: Le Kou, affluent de la Volta Noire, à 30 km au Nord de Bobo-Dioulasso , République de Haute Volta. [ Burkina Faso]. Holotype: RMCA 141464 View Materials .
Distribution: 505, 507, 508, 509, 511, 514, 515, 516, 520, 535 (Gosse et al., 2003; RMCA fish collection).
Nannocharax brevis Boulenger, 1902
Nannocharax brevis Boulenger, 1902c:27 , Pl. 8 (fig. 2). Type locality: L’Ubangi à Banzyville, Congo. [ Democratic Republic of the Congo]. Syntypes: BMNH 1901.12 . 26.21 (1), RMCA 1282 View Materials (1) , 1282A (1).
Distribution: 535, 536, 537, 539, 540, 545, 546, 549 (RMCA fish collection; Boulenger, 1909a; David, Poll, 1937; Depasse, 1956; Lambert, 1961; Gosse, 1968; Gosse, 1963; Poll, 1976; Mbimbi Mayi Munene et al., 2021).
Nannocharax dageti Jerep, Vari, Vreven, 2014
Nannocharax dageti Jerep, Vari, Vreven, 2014:362 View in CoL , figs. 1–2. Type locality: Kando GoogleMaps , 10°48’ S 26°13’ E, Democratic Republic of the Congo. Holotype: RMCA 88846 View Materials .
Distribution: 544, 545, 546, 555, 569 (Jerep et al., 2014; Bragança et al., 2020).
Nannocharax elongatus Boulenger, 1900
Nannocharax elongatus Boulenger, 1900b:131 , Pl. 48 (fig. 3). Type locality: Haut-Congo , entre Nouvelle-Anvers et Stanley Falls, État indépendant du Congo. [ Democratic Republic of the Congo]. Syntypes: BMNH 1900.12.13.26–27 (2), RMCA 263 View Materials to 268 View Materials (1,1,1,1,1,1).
Distribution: 535, 536, 537, 539, 540, 545, 546 (RMCA fish collection; Depasse, 1956; Boulenger, 1909a; Boulenger, 1920b; Pellegrin, 1928b; David, Poll, 1937; Gosse, 1963; Gosse, 1968; Monsembula Iyaba, Stiassny, 2013; Katemo Manda et al., 2021).
Nannocharax fasciatus Günther, 1867
Nannocharax fasciatus Günther, 1867:112 View in CoL [4], Pl. 3 (fig. A). Type locality: Gaboon. Holotype: BMNH 1867.5.3.35.
Distribution: 505, 506, 507, 508, 509, 511, 512, 514, 515, 516, 517, 518, 520, 534 (Daget, 1978; Vari , 2007; Gosse et al., 2003; RMCA fish collection).
Nannocharax fasciolaris Nichols, Boulton, 1927
Nannocharax fasciolaris Nichols, Boulton, 1927:3 View in CoL , fig. 1. Type locality: Capelongo , Angola. Holotype: AMNH 8683 View Materials .
Distribution: 555 (Daget, Gosse, 1984).
Nannocharax gobioides Roman, 1966
Nannocharax gobioides Roman, 1966:78 View in CoL , Pl. 5 (figs. 9–10). Type locality: Volta Noire , à 47 km au Nord de Bobo-Dioulasso, Haute Volta. [ Burkina Faso]. Holotype: RMCA 141442 View Materials .
Distribution: 516 (Roman, 1966).
Remarks: Nannocharax gobioides is treated as a synonym of Nannocharax fasciatus in Gosse, Coenen (1990), who refer to Coenen (1990: unpublished manuscript) for the synonymy, and in Lévêque et al. (1991), who refer to presumably the same publication from Coenen as “Coenen (in prep.)”. This synonymy is also given in Gosse et al. (2003), without the reference to Coenen (1990: unpublished manuscript), but this seems merely a repetition of the data in Gosse, Coenen (1990). As Coenen (1990) refers to a non-published draft manuscript (Erik Coenen, pers. comm.), which is presumably the same as Coenen (in prep.), any taxonomical change refering to this publication seems unsubstantiated.
Nannocharax gracilis Poll, 1939
Nannocharax gracilis Poll, 1939:30 , fig. 14. Type locality: Léopoldville , Congo Belge. [ Democratic Republic of the Congo]. Syntypes: RMCA 39708 View Materials (1), 39866–76 (11), 39877–919 (43), 43979 (1).
Distribution: 535, 540, 546, 548 (Poll, 1939; Poll, 1959; Lambert, 1961; Gosse, 1963).
Nannocharax hadros Katemo Manda, Snoeks, Chocha Manda, Vreven, 2021
Nannocharax hadros Katemo Manda, Snoeks, Chocha Manda, Vreven, 2021:319 , fig. 1. Type locality: Democratic Republic of the Congo: Upemba National Park GoogleMaps , Kalule Nord River, bridge on the road Lubudi-Luena, 9°41’36”S 25°50’52”E, 790 m a.s.l. Holotype: RMCA 2016-025 View Materials - P-0004 .
Distribution: 545 (Katemo Manda et al., 2021).
Nannocharax hastatus Jerep, Vari, 2014
Nannocharax hastatus Jerep, Vari, 2014:44 , fig. 1. Type locality: Republic of the Congo, Cuvette-Ouest GoogleMaps , Likouala River GoogleMaps drainage, small canal around island in Lékoli River, Odzala National Park, 0°37’14”N, 14°55’5.9”E. Holotype: CUMV 89072 View Materials .
Distribution: 534, 535, 538, 548 (Jerep, Vari , 2014).
Nannocharax hollyi Fowler, 1936
Nannocharax hollyi Fowler, 1936:271 View in CoL , fig. 21. Type locality: Fort Sibut , Ubangi-Shari. [ Central African Republic]. Holotype: ANSP 65558 View Materials .
Distribution: 535 (Fowler, 1936; Daget, Gosse, 1984).
Nannocharax intermedius Boulenger, 1903
Nannocharax intermedius Boulenger, 1903a:437 View in CoL . Type locality: Kribi River , southern Cameroon. Syntypes: BMNH 1903.7.28.144–147 (4).
Distribution: 505, 533, 534 ( Vari , 2007; RMCA fish collection).
Nannocharax latifasciatus Coenen, Teugels, 1989
Nannocharax latifasciatus Coenen, Teugels, 1989:312 , fig. 1. Type locality: Affluent GoogleMaps of the New Calabar River, 3 km south of Isiokpo (4°57’N, 6°53’E), Nigeria. Holotype: RMCA 88-12 View Materials - P-1 .
Distribution: 506, 518 (Gosse et al., 2003; Vari , 2007).
Nannocharax lineomaculatus Blache, Miton, 1960
Nannocharax lineomaculatus Blache, Miton, 1960:105 . Type locality: Gamsaye (Logone Inf.), Chaoué ( Bas Chari ), Logone Birni (Logone Inf.), Fort-Lamy et Zymado (Logone Inf.). [Gamsaye (lower Logone River) and Fort-Lamy are in Republic of Chad, while Chaoué (= Chawe, on the lower Chari River), Logone- Birni and Zymado (both lower Logone River) seem to be in Cameroon, on or near the border with Chad]. Syntypes: MNHN 1959-0230 View Materials (6); CEPFL 716–723 (8), CEPFL 1369–81 (13). [In the original description the MNHN specimens are cited as types and the CEPFL specimens as paratypes. They are all to be treated as syntypes (See ICZN, 1999: Art. 72.1 and 73.2)].
Distribution: 505, 507, 508, 520 (Gosse et al., 2003).
Nannocharax lineostriatus (Poll, 1967)
Hemigrammocharax lineostriatus Poll, 1967:123 View in CoL , fig. 52. Type locality: Alto Cuílo, mare des rives du Cuílo, Angola. Holotype: MD 5342 .
Distribution: 546 (Poll, 1967. Skelton, 2019 also identified specimens from 551 and 555 as Nannocharax lineostriatus , but Bragança et al., 2020 use Nannocharax cf. lineostriatus for specimens from these areas.).
Nannocharax luapulae Boulenger, 1915
Nannocharax luapulae Boulenger, 1915:164 View in CoL . Type locality: Rivière Luapula , à Kasenga. [ Democratic Republic of the Congo]. Holotype: RMCA 12125 View Materials .
Distribution: 540, 544, 545 (Boulenger, 1915; David, Poll, 1937; Poll, 1976).
Nannocharax machadoi (Poll, 1967)
Hemigrammocharax machadoi Poll, 1967:125 View in CoL , fig. 53. Type locality: Riv. Longa, afflt. Luena , village Nharicumbi, Angola. Holotype: MD 5579 .
Distribution: 555, 556, 557, 558, 561, 569 (Skelton, 2001).
Nannocharax macropterus Pellegrin, 1926
Nannocharax macropterus Pellegrin, 1926:163 . Type locality: Mongende , Luebo ( Kasai ), Kamaiembi ( Kasai ) and Tschikapa (Kasai), Congo belge. [ Democratic Republic of the Congo]. Syntypes: MNHN 1925-0313 View Materials to 0315 View Materials (4); RMCA 19698 View Materials (2), 19703 (1), 19715 (1), 19616–17 (2). [(1) In the original description a total of 38 type specimens are mentioned, including 31 from Tschikapa. Pellegrin (1928b) described this species as new again with the same data as in Pellegrin (1926), but mentioned only 3 specimens from Tshikapa; presumably the count of 31 type specimens from Tshikapa in Pellegrin (1926) was a typographic error, so that the actual number of type specimens is 10. However, counting the available syntypes in collection, there are 3 from Mongende (3 reported), 4 from Luebo (only 2 reported), 1 from Kamaiembi (2 reported) and 2 from Tshikapa (3 reported). The total number of syntypes in museum collections matches the 10 types mentioned in Pellegrin (1928b), but the number of specimens for the different type localities does not. (2) RMCA 19698 View Materials was verified as containing 2 specimens. (3) the MNHN online collection database reports MNHN 1958- 0086 View Materials as a syntype, but this specimen is however not collected at one of the type localities and thus excluded from the type series].
Distribution: 535, 536, 537, 540, 546, 548, 549, 555, 556, 557, 569 (RMCA fish collection; Pellegrin, 1926; David, Poll, 1937; Poll, 1959; Lambert, 1961; Poll, 1967; Skelton, 2001; Decru et al., 2017).
Nannocharax maculicauda Vari, Géry, 1981 View in CoL
Nannocharax maculicauda Vari, Géry, 1981:1076 View in CoL , figs. 1–4. Type locality: Still GoogleMaps side arm of the upper Ivindo River GoogleMaps opposite Bourassié GoogleMaps , Gabon, at the juncture of the Djouah and Karouga Rivers (approx. 1°20’N, 13°12’E). Holotype: MNHN 1981-0608 View Materials .
Distribution: 532, 533 ( Vari , 2007; Jerep, Vari , 2014).
Nannocharax micros Fowler, 1936
Nannocharax micros Fowler, 1936:268 View in CoL , fig. 19. Type locality: Thirty kilometers east of Kribi, Cameroons [ Cameroon]. Holotype: ANSP 65556 View Materials .
Distribution: 533 (Daget, Gosse, 1984).
Nannocharax minutus Worthington, 1933 View in CoL
Nannocharax minutus Worthington, 1933:40 View in CoL . Type locality: Swamp near the River Luombo, Luapula system, Northern Rhodesia. [ Zambia]. Syntypes: BMNH 1932.12.16.45–54 (10).
Distribution: 544, 555 (RMCA fish collection; Worthington, 1933; Van Steenberge et al., 2014; Bragança et al., 2020).
Nannocharax monardi (Pellegrin, 1936)
Hemigrammocharax monardi Pellegrin, 1936:49 View in CoL . Type locality: Kukulakazé (affluent du Kului) et Vila da Ponte (Cubango), Angola. Syntypes: MNHN 1936-0073 View Materials (2, apparently lost).
Distribution: 555 (Pellegrin, 1936).
Nannocharax multifasciatus Boulenger, 1923
Nannocharax multifasciatus Boulenger, 1923:437 View in CoL . Type locality: Sesheke , R. Zambezi, Northern Rhodesia. [ Zambia]. Holotype: SAIAB 135308 View Materials .
Distichodus stigmaturus Fowler, 1935:260 View in CoL , figs. 6–7. Type locality: Chobe River, below rapids, three miles from Kasane. Holotype: SAIAB [formerly RUSI] 30005.
Distribution: 544, 552, 553, 555, 556, 557, 569 (Daget, Gosse, 1984; van der Waal, Skelton, 1994; Skelton, 1994; Skelton, 2001).
Nannocharax niloticus (Joannis, 1835)
Coregonus niloticus Joannis, 1835 :no page number, Pl. 7. Type locality: Thèbes. [ Egypt]. Holotype: Type lost.
Nannocharax niloticus var. lateralis Werner, 1906:1130 View in CoL , Pl. 3 (figs. 11, 11a). Type locality: Bahr-el-Gebel bei Mongalla. [ South Sudan]. No types known.
Nannocharax niloticus tchadensis Blache, Miton, 1961:396 View in CoL . Type locality: Logone moyen, zones inondées vers Pouss et Gamsaye; Mayo Kebbi, zones des rapides en amont des Chutes Gauthiot; Logone moyen à Gabri Ngolo. [Middle Logone River, floodplain near Pouss and Gamsaye (border Republic of Chad- Cameroon); Mayo Kebbi, rapids zone upstream of the Gauthiot Falls ( Republic of Chad); middle Logone at Gabri Ngolo ( Republic of Chad)]. Syntypes: MNHN 1959-0246 (2), 1959-0347 (3), 1959-0248 (orig. 25, now 15), 1988- 1111 (10). [(1) The total number of syntypes adds up to a total of 30 specimens, while the original description seems to be based on 36 specimens. (2) the original description mentioned both “types” and paratypes, all to be treated as syntypes (See ICZN, 1999: Art. 72.1 and 73.2)]. Gosse, Coenen (1990) put N. niloticus View in CoL tchadensis in synonymy with both N. occidentalis View in CoL (partim) and N. lineomaculatus View in CoL (partim), for which they refer to Coenen (1990: unpublished manuscript). The partim synonymy with both N. occidentalis View in CoL and N. lineomaculatus View in CoL is also given in Gosse et al. (2003), but this seems merely a repetition of the data in Gosse, Coenen (1990). As Coenen (1990) is a non-published draft manuscript (Erik Coenen, pers. comm.), any taxonomical change referring to this publication seems unsubstantiated. Nannocharax niloticus View in CoL tchadensis is considered here to be a synonym of N. niloticus View in CoL based on Daget, Gosse (1984).
Distribution: 522, 523, 524 (Boulenger, 1907c; Lévêque et al., 1991).
Nannocharax occidentalis Daget, 1959
Nannocharax niloticus occidentalis Daget, 1959:669 . Type locality: Markala, passe à poissons; Dabola, chutes du Tinkisso; Bourem, mission Augiéras-Draper. [ Markala , fish ladder ( Mali); Dabola , Tinkisso falls ( Guinea); Bourem , Augiéras-Draper mission ( Mali)]. Syntypes: MNHN 1954-0010 View Materials (3), 1928-0235 (3), 1960- 0414 (3, now 2), 1960-0415 (3), 1988-1110 (1). [The MNHN online collection database reports 3 specimens for MNHN 1960-0414 View Materials ; this is probably an error, as a single specimen was removed and relabelled as MNHN 1988-1110 View Materials . Counting only 2 specimens for MNHN 1960-0414 View Materials brings the total number of syntypes on 12, as reported in the original description]. Replacement for Nannocharax niloticus gracilis Daget, 1954 preoccupied by Nannocharax gracilis Poll, 1939 .
Nannocharax niloticus gracilis Daget, 1954:163 , fig. 51. Type locality: Markala, passe à poissons; Dabola, chutes du Tinkisso; Bourem, mission Augiéras-Draper. [Markala, fish ladder ( Mali); Dabola, Tinkisso falls ( Guinea); Bourem, Augiéras-Draper mission ( Mali)]. Syntypes: MNHN 1954-0010 (3), 1928-0235 (3), 1960-0414 (orig. 3, now 2), 1960-0415 (3), 1988-1110 (1). Preoccupied by Nannocharax gracilis Poll, 1939 , replaced by Nannocharax niloticus occidentalis Daget, 1959 (currently valid as Nannocharax occidentalis Daget, 1959 ). Gosse, Coenen (1990) put N. niloticus gracilis in synonymy with N. occidentalis (partim) and N. lineomaculatus (partim), for which they refer to Coenen (1990: unpublished manuscript). This partim synonymy with both N. occidentalis and N. lineomaculatus is also given in Gosse et al. (2003), but this seems merely a repetition of the data in Gosse, Coenen (1990). As Coenen (1990) is a non-published draft manuscript (Erik Coenen, pers. comm.), any taxonomical change referring to this publication seems unsubstantiated. Nannocharax niloticus gracilis is considered here to be a synonym of N. occidentalis only, following Daget, Gosse (1984) and Levêque et al. (1991).
Distribution: 505, 506, 507, 508, 518, 520 (Gosse et al., 2003).
Nannocharax ocellicauda Boulenger, 1907
Nannocharax ocellicauda Boulenger, 1907a:485 View in CoL . Type locality: Ja River , South Cameroon. Syntypes: BMNH 1908.5.30.121–122 (2).
Distribution: 533, 534 ( Vari , 2007; Jerep, Vari , 2014).
Nannocharax ogoensis Pellegrin, 1911
Nannocharax ogoensis Pellegrin, 1911:180 View in CoL . Type locality: Franceville (Ogôoué), Congo français. [ Gabon]. Holotype: MNHN 1886-0397 View Materials .
Distribution: 532 (Daget, Gosse, 1984).
Nannocharax parvus Pellegrin, 1906
Nannocharax parvus Pellegrin, 1906c:469 View in CoL . Type locality: Ngomo (Ogôoué), Gabon. Syntypes: BMNH 1907.4 . 27.2 (1), MNHN 1906-0194 View Materials (4).
Nannocharax parvus var. maculata Pellegrin, 1924a:285 View in CoL . Type locality: Fleuve Niari , Gabon. Syntypes: MNHN 1924-0049 View Materials to 0051 View Materials (3).
Distribution: 532, 550 ( Vari , 2007; RMCA fish collection; Dunz, Schliewen, 2009; Wamuini Lunkayilakio et al., 2010).
Nannocharax procatopus Boulenger, 1920
Nannocharax procatopus Boulenger, 1920b:20 , fig. 9. Type locality: Bosabangi et Poko, Congo Belge. [ Democratic Republic of the Congo]. Syntypes: BMNH 1919.9 . 10.177 (1), 1919.9.10.178 (1); RMCA 7156–57 View Materials (2) , 7165 (1).
Distribution: 534, 536, 537, 539, 540, 546 (RMCA fish collection; BMNH fish collection; Boulenger, 1920b; Pellegrin, 1928b; Poll, 1967; Decru et al., 2017; Dunz, Schliewen, 2009).
Nannocharax pteron Fowler, 1936
Nannocharax pteron Fowler, 1936:272 View in CoL , fig. 22. Type locality: Fort Sibut , Ubangi-Shari. [ Central African Republic]. Holotype: ANSP 65560 View Materials .
Distribution: 535, 537 (Fowler, 1936; Daget, Gosse, 1984; Decru et al., 2017).
Nannocharax reidi Vari, Ferraris, 2004 View in CoL
Nannocharax reidi Vari, Ferraris, 2004:552 View in CoL , fig. 1. Type locality: Cameroon, Cross River GoogleMaps system, collecting points on southern Munaya River GoogleMaps draining northern Korup GoogleMaps , on Basep River at junction with Munaya River (5°49’30”N, 9°03’30”E). Holotype: USNM 304046 View Materials .
Distribution: 518 ( Vari , 2007).
Nannocharax rubensteini (Jerep, Vari, 2013)
Hemigrammocharax rubensteini Jerep, Vari, 2013:31 , figs. 1–3. Type locality: Republic of the Congo, Cuvette-Ouest GoogleMaps , Likouala River GoogleMaps drainage, Lékoli River GoogleMaps at Mboko dock, Odzala National Park, 0°37’03”N, 14°54’27”E. Holotype: CUMV 89502 View Materials .
Distribution: 534 (Jerep, Vari , 2013).
Nannocharax rubrolabiatus Van den Bergh, Teugels, Coenen, Ollevier, 1995 View in CoL
Nannocharax rubrolabiatus Van den Bergh, Teugels, Coenen, Ollevier, 1995:350 View in CoL , figs. 2–3. Type locality: Cameroon, Noun River GoogleMaps , affluent of Mbam River GoogleMaps , Sanaga GoogleMaps basin at the bridge on the road from Bafoussam to Foumbot; 5°29’N, 10°31’E. Holotype: RMCA 94-74 View Materials - P-1 .
Distribution: 505 ( Vari , 2007).
Nannocharax schoutedeni Poll, 1939
Nannocharax schoutedeni Poll, 1939:29 , fig. 13. Type locality: Léopoldville , Congo Belge. [ Democratic Republic of the Congo]. Syntypes: (119) RMCA 39705–07 View Materials (3), 39920–32 (13), 39933–40029 (98), 43883–86 (4), 44016 (1). [Poll (1939) mentioned 16 types and 101 cotypes which adds up to 117 syntypes. However, the number of syntypes in the RMCA collection is 119 ( RMCA 39933–40029 View Materials actually contains 98 specimens), all from the type locality and in agreement with the collector and collection date given in the original description].
Distribution: 535, 537, 546, 548 (RMCA fish collection; Gosse, 1963; Gosse, 1968; Dunz, Schliewen, 2009; Monsembula Iyaba, Stiassny, 2013; Stiassny et al., 2021).
Nannocharax seyboldi Schultz, 1942
Nannocharax seyboldi Schultz , 1942313, Pl. 35 (fig. 1). Type locality: Bellyella , Liberia. Holotype: USNM 118757 View Materials .
Distribution: 512 (Schultz, 1942; Jerep et al., 2014; Jerep, Vari , 2014).
Remarks: Nannocharax seyboldi is treated as a synonym of Nannocharax fasciatus in Gosse, Coenen (1990), who refer to Coenen (1990: unpublished manuscript) for the synonymy. This synonymy is also given in Gosse et al. (2003), without the reference to Coenen (1990: unpublished manuscript), but this seems merely a repetition of the data in Gosse, Coenen (1990). As Coenen (1990) is a non-published draft manuscript (Erik Coenen, pers. comm.), any taxonomical change referring to this publication seems unsubstantiated. Jerep et al. (2014) and Jerep, Vari (2014) included the types of N. seyboldi in a list with examined material, but did not mention the taxonomic status of the species.
Nannocharax signifer Moritz, 2010
Nannocharax signifer Moritz, 2010:290 , figs. 1–2. Type locality: Benin: Forêt d’Agrimè GoogleMaps , about 7 kilometers east of Zogbodome, Hlan River, 7°03’58”N, 2°10’56”E. Holotype: BMNH 2009.3 .2.1.
Distribution: 517 (Moritz, 2010).
Nannocharax taenia Boulenger, 1902
Nannocharax taenia Boulenger, 1902c:28 View in CoL , Pl. 8 (fig. 3). Type locality: Banzyville , Congo. [ Democratic Republic of the Congo]. Syntypes: BMNH 1901.12 . 26.22 (1), RMCA 1283 View Materials (1).
Distribution: 534, 535, 536, 537, 540, 546 (RMCA fish collection; Nichols, Griscom, 1917; David, Poll, 1937; Gosse, 1963; Matthes, 1964b; Dunz, Schliewen, 2009; Ibala Zamba et al., 2019).
Nannocharax uniocellatus (Pellegrin, 1926)
Microdistichodus uniocellatus Pellegrin, 1926:162 . Type locality: Mongende et Luebo (Kasai), Congo Belge. [ Democratic Republic of the Congo]. Syntypes: MNHN 1925-0311 View Materials (5), 1925-0312 (1); RMCA 19704–08 View Materials (5) , 19718–19 (2).
Distribution: 534, 535, 537, 540, 546, 548 (RMCA fish collection; AMNH fish collection; Monsembula Iyaba, Stiassny, 2013; Jerep et al., 2014).
Nannocharax usongo Dunz, Schliewen, 2009
Nannocharax usongo Dunz, Schliewen, 2009:10 View in CoL , figs. 5a–c, 6. Type locality: Cameroon, Cross River GoogleMaps at Mamfé, 5°46’0.00”N, 9°18’38.00”E Holotype: ZSM 35732 View Materials .
Distribution: 506, 518 (Dunz, Schliewen, 2009).
Nannocharax wittei (Poll, 1933)
Hemigrammocharax wittei Poll, 1933:122 , fig. 8. Type locality: Kando (près de Tenke): Riv. Kando, Congo Belge. [ Democratic Republic of the Congo]. Syntypes: RMCA 34903–18 View Materials (16), 34919–51 (33), 36247–311 (65).
Distribution: 544, 545, 546, 555 (RMCA fish collection; Poll, 1967).
Nannocharax zebra Dunz, Schliewen, 2009
Nannocharax zebra Dunz, Schliewen, 2009:5 View in CoL , figs. 3a, b, 4. Type locality: Cameroon, Cross River GoogleMaps at Mamfé, 5°46’0.00”N, 9°18’38.00”E. Holotype: ZSM 35730 View Materials .
Distribution: 506, 518 (Dunz, Schliewen, 2009).
Neolebias Steindachner, 1894:78 . Type species: Neolebias unifasciatus Steindachner, 1894 . Type by monotypy. Gender: Masculine.
Rhabdaethiops Fowler, 1936:264 . Type species: Nannaethiops tritaeniatus Boulenger, 1913 . Type by original designation. Gender: Masculine. Additional comments: Originally proposed as a subgenus of Nannaethiops .
Micraethiops Daget, 1965:20 . Type species: Neolebias ansorgii Boulenger, 1912 . Type by original designation. Gender: Masculine. Additional comments: Originally proposed as a subgenus of Neolebias .
Neolebias ansorgii Boulenger, 1912 Neolebias ansorgii Boulenger, 1912b:8 , Pl. 17 (fig. 3). Type locality: Marécages près de la Luculla (Congo belge) et de la Luali à Lundo (Mayombé Portugais). [Syntype locality: swamps near the Luculla River (Chiloango River basin in Belgian Congo = Democratic Republic of the Congo) and the Luali River at Lundo ( Chiloango River basin in “ Portuguese Mayombe ”, Cabinda Province of Angola). Lectotype locality: marshes near riv. Luculla , Mayumbe ( Chiloango River basin in Democratic Republic of the Congo)]. Lectotype: BMNH 1912.4 .1.276. [Designated by Poll, Gosse (1963)].
Neolebias landgrafi Ahl, 1928b:206 View in CoL . Type locality: Kamerun. [ Cameroon]. Syntypes: ZMB 31686–87 View Materials (2, 5).
Nannaethiops geisleri Hoedeman, 1956b:260 View in CoL , fig. Type locality: Congo-Gebiet. [Holotype locality was not specified (“Congo-Gebiet”), while the 3 paratypes are either from near Boma (“ Umgebung Boma ”) as given in the species description, or from near Leopoldville (“ Umgebung von Leopoldville ”) as given in the publication’s introduction]. Holotype: ZMA 100053 View Materials .
Distribution: 506, 517, 532, 550 (Gosse et al., 2003; Vari , 2007; Poll, Gosse, 1963a).
Neolebias axelrodi Poll, Gosse, 1963
Neolebias axelrodi Poll, Gosse, 1963a:31 View in CoL , Pl. 2 (fig. 3). Type locality: Source à environ 30 miles de Lagos le long de la nouvelle route. [ Nigeria]. Holotype: BMNH 1970.12 .14.200.
Distribution: 506, 517 (Gosse et al., 2003).
Neolebias gossei (Poll, Lambert, 1964)
Congocharax gossei Poll, Lambert, 1964:337 View in CoL , figure. Type locality: Lambaréné-Mouila ( PK9 ), Ogooué, Gabon. Holotype: RMCA 142822 View Materials .
Distribution: 532 ( Vari , 2007).
Neolebias gracilis Matthes, 1964
Neolebias gracilis Matthes, 1964b:69 View in CoL , Pl. 3a. Type locality: Riv. Boteli (région d’Ikela ), Congo. [ Democratic Republic of the Congo]. Holotype: RMCA 138900 View Materials .
Distribution: 537 (Monsembula Iyaba, Stiassny, 2013; Matthes, 1964b).
Neolebias kerguennae Daget, 1980
Neolebias kerguennae Daget, 1980:33 View in CoL , fig. 1. Type locality: Ruisseau à 29 km à l’est de Libreville, Gabon. Holotype: MNHN 1980-1299 View Materials .
Distribution: 532 ( Vari , 2007).
Neolebias lozii Winemiller, Kelso-Winemiller, 1993 View in CoL
Neolebias lozii Winemiller, Kelso-Winemiller, 1993:112 View in CoL , fig. 1. Type locality: Kataba Creek GoogleMaps , Upper Zambezi drainage, at the Mongu-Senanga highway, Western Province, Zambia, 15°30’30”S, 23°16’0”E. Holotype: USNM 320688 View Materials .
Distribution: 556 ( Winemiller, Kelso-Winemiller, 1993).
Neolebias philippei Poll, Gosse, 1963
Neolebias philippei Poll, Gosse, 1963a:30 View in CoL , Pls. 1 (fig. 4), 2 (fig. 2). Type locality: Boende , riv. Tshuapa. [ Democratic Republic of the Congo]. Holotype: RMCA 98971 View Materials .
Distribution: 537, 540, 546 (Poll, Gosse, 1963a; Daget, Gosse, 1984; Decru et al., 2017; Stiassny et al., 2021a).
Neolebias powelli Teugels, Roberts, 1990
Neolebias powelli Teugels, Roberts, 1990:61 , figs. 1–4. Type locality: Sombreiro river, Niger delta , at Rumuekpe, Nigeria (4°57’N, 6°40’E). Holotype: RMCA 87-24-P-135.
Distribution: 506 (Gosse et al., 2003).
Neolebias spilotaenia Boulenger, 1912
Neolebias spilotaenia Boulenger, 1912b:9 , Pl. 17 (fig. 4 [not 3]). Type locality: Lebuzi à Kuka Muno (Congo Belge) et Luali à Lundo ( Congo Portugais ). [Lebuzi River at Kuka Muno (Belgian Congo = Democratic Republic of the Congo) and Luali River at Lundo (Portuguese Congo = Cabinda Province of Angola), both in the Chiloango River basin. Lectotype locality is “Mayumbe Kuka Muno, Lebuzi River”. Democratic Republic of Congo]. Lectotype: BMNH 1912.4 .1.282. [Designated by Poll, Gosse (1963a)].
Distribution: 532 ( Vari , 2007).
Neolebias trewavasae Poll, Gosse, 1963
Neolebias trewavasae Poll, Gosse, 1963a:27 View in CoL , Pl. 2 (fig. 1). Type locality: Yaekama , étang. [ Democratic Republic of the Congo]. Holotype: RMCA 137767 View Materials .
Distribution: 505, 522, 532, 533, 534, 535, 537, 539, 540, 541, 546 (Sandon, 1950; Poll, Gosse, 1963a; Vari , 2007; Decru et al., 2017; RMCA fish collection).
Neolebias trilineatus Boulenger, 1899
Neolebias trilineatus Boulenger, 1899:114 View in CoL . Type locality: Dolo (Stanley-Pool), Congo. [ Democratic Republic of the Congo]. Lectotype: RMCA 729 View Materials . [Designated by Poll, Gosse (1963a)].
Nannaethiops tritaeniatus Boulenger, 1913:157 View in CoL . Type locality: Dungu, district du Haut-Uélé, Congo belge. [ Democratic Republic of the Congo]. Syntypes: BMNH 1912.12 . 6.2 (1); RMCA 1796–1797 View Materials (2) , 1815 (1).
Distribution: 534, 535, 536, 537, 540, 542, 546, 548 (RMCA fish collection; AMNH fish collection; Monsembula Iyaba, Stiassny, 2013).
Neolebias unifasciatus Steindachner, 1894
Neolebias unifasciatus Steindachner, 1894:78 View in CoL , Pl. 4 (fig. 3). Type locality: Robertsport , Liberia. Lectotype: RMNH 24926 View Materials . [Designated by Poll, Gosse (1963)].
Neolebias unitaeniatus Boulenger, 1899c:114 (footnote). Type locality: Robertsport , Liberia. Lectotype: RMNH 24926 View Materials . Unneeded replacement name for Neolebias unifasciatus Steindachner, 1894 View in CoL .
Neolebias univittatus Boulenger, 1899c:114 (footnote). Type locality: Robertsport , Liberia. Lectotype: RMNH 24926 View Materials . Unneeded replacement name for Neolebias unifasciatus Steindachner, 1894 .
Nannaethiops angustolinea Nichols, 1923:1 View in CoL , fig. 1. Type locality: Fort Crampel , Gribingui, West Africa, French Equatorial Africa. [ Central African Republic]. Holotype: AMNH 8106 View Materials .
Nannocharax shariensis Fowler, 1949b:248 View in CoL , figs. 24–27. Type locality: Ubangui-Shari, near N’délé, French Equatorial Africa. [ Central African Republic]. Holotype: ANSP 71888 View Materials .
Distribution: 505, 506, 507, 508, 509, 511, 512, 514, 515, 516, 517, 520, 532, 533, 534, 535 (Gosse et al., 2003; Vari , 2007; RMCA fish collection; Vivien, 1991; Daget, Gosse, 1984).
PARADISTICHODUS Pellegrin, 1922
Paradistichodus Pellegrin, 1922:70 . Type species: Paradistichodus elegans Pellegrin, 1922 . Type by original designation. Gender: Masculine.
Paradistichodus dimidiatus (Pellegrin, 1904)
Nannocharax dimidiatus Pellegrin, 1904a:220 View in CoL . Type locality: Casamance, à la hauteur de Kolibantan, Guinée portugaise. [ Guinea-Bissau]. Holotype: MNHN 1904- 0055 View Materials .
Distichodus ansorgii Boulenger, 1911a:373 View in CoL . Type locality: Geba and Culufi Rivers, Portuguese Guinea. [ Guinea-Bissau]. Syntypes: BMNH 1912.4.1.65–68 (4), 1912.4.1.69–74 (6), 1912.4.1.75–77 (3); CAS 60707 (1) ; RMCA 7074 View Materials (1) ; ZMB 18444 View Materials (6) Culufi River .
Paradistichodus elegans Pellegrin, 1922:72 View in CoL . Type locality: Bossangoa (Ouham). [ Central African Republic]. Syntypes: MNHN 1921-0216 View Materials , MNHN 1921-0217 View Materials . [two syntypes, both are apparently lost] .
Distribution: 505, 506, 507, 508, 509, 511, 516, 520 (Gosse et al., 2003).
Paraphago Boulenger, 1899:76 . Type species: Paraphago rostratus Boulenger, 1899 . Type by monotypy. Gender: Masculine.
Paraphago rostratus Boulenger, 1899
Paraphago rostratus Boulenger, 1899b:76 , Pl. 34 (fig. 2). Type locality: Kutu (Lac Léopold II), Congo. [ Democratic Republic of the Congo]. Syntypes: BMNH 1899.9 . 26.81 (1), RMCA 716–717 View Materials (2).
Distribution: 547 (Boulenger, 1909a; Daget, Gosse, 1984).
Phago Günther, 1865b:209 . Type species: Phago loricatus Günther, 1865 . Type by monotypy. Gender: Masculine.
Phago boulengeri Schilthuis, 1891
Phago boulengeri Schilthuis, 1891:90 . Type locality: Not specified. [Collections studied in this publication originated from “Boma, lower Congo, and for the greater part from Kinshassa, Stanley Pool, upper Congo, near Brazzaville” in Democratic Republic of the Congo. Apparently only the species not collected at Kinshassa have their locality specifically stated, so that the holotype of P. boulengeri probably also originated from the Stanley Pool in Kinshassa]. Holotype: BMNH 1899.9 .6.12.
Distribution: 534, 535, 536, 537, 538, 540, 545, 546, 547, 548 (RMCA fish collection; AMNH fish collection; Nichols, Griscom, 1917; Poll, 1933; David, Poll, 1937; Poll, 1959; Gosse, 1963; Matthes, 1964b; Gosse, 1968).
Phago intermedius Boulenger, 1899
Phago intermedius Boulenger, 1899b:77 View in CoL , Pl. 34 (fig. 3). Type locality: Léopoldville , Etat du Congo. [ Democratic Republic of the Congo]. Holotype: RMCA 123 View Materials .
Phago fulica Pellegrin, 1922:69 View in CoL . Type locality: Bangui ( Oubanghi ). [ Central African Republic]. Holotype: MNHN 1921-0374 View Materials .
Distribution: 534, 535, 536, 537, 540, 545, 548 (RMCA fish collection; AMNH fish collection; Nichols, Griscom, 1917; Poll, 1933; David, Poll, 1937; Poll, 1959; Gosse, 1963; Matthes, 1964b; Gosse, 1968).
Phago loricatus Günther, 1865b:210 [2], Pl. 5. Type locality: Western coast of Africa. [The original collection register of the BMNH (now NHMUK), of which a (digital) copy is provided in the online database of the BMNH fish collection, records “ W. Africa ” as origin of the specimen. This was recorded as “Niger ” for both “locality” and “water body” in the respective fields in the online database. Until more information on or a source for this change is available, the original type locality should be used]. Holotype: BMNH 1865.1 .20.15.
Phago maculatus Ahl, 1922:23 , figure. Type locality: Unterer Niger . Holotype: ZMB 20558 View Materials .
Distribution: 505, 506, 517, 518 (Gosse et al., 2003; Vari , 2007).
Xenocharax Günther, 1867:114 . Type species: Xenocharax spilurus Günther, 1867 . Type by monotypy. Gender: Masculine.
Xenocharax crassus Pellegrin, 1900
Xenocharax crassus Pellegrin, 1900:100 View in CoL . Type locality: Leketi (Alima moyen), Congo français. [ Republic of the Congo]. Holotype: MNHN 1886-0384 View Materials .
Distribution: 534, 535, 537, 538, 546, 547, 548 (Ibala Zamba et al., 2016; RMCA fish collection; Monsembula Iyaba, Stiassny, 2016).
Remarks: revalidated by Ibala Zamba et al. (2016).
Xenocharax spilurus Günther, 1867
Xenocharax spilurus Günther, 1867:114 View in CoL [6], Pl. 3 (fig. B). Type locality: Gaboon [ Gabon]. Lectotype: BMNH 1867.5 .3.33. [Designated by Ibala Zamba et al. (2016)].
Distribution: 532, 533 ( Vari , 2007; RMCA fish collection).
No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.
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DISTICHODONTIDAE Günther, 1864
Toledo-Piza, Mônica, Baena, Eduardo G., Dagosta, Fernando C. P., Menezes, Naércio A., Ândrade, Marcelo, Benini, Ricardo C., Bertaco, Vinicius A., Birindelli, José Luís O., Boden, Gert, Buckup, Paulo A., Camelier, Priscila, Carvalho, Fernando R., Castro, Ricardo M. C., Chuctaya, Junior, Decru, Eva, Derijst, Eddy, Dillman, Casey B., Ferreira, Katiane M., Merxem, Dimitri G., Giovannetti, Victor, Hirschmann, Alice, Jégu, Michel, Jerep, Fernando C., Langeani, Francisco, Lima, Flávio C. T., Lucena, Carlos A. S., Lucena, Zilda Margarete S., Malabarba, Luiz R., Malabarba, Maria Cláudia S. L., Marinho, Manoela M. F., Mathubara, Kleber, Mattox, George M. T., Melo, Bruno F., Moelants, Tuur, Moreira, Cristiano R., Musschoot, Tobias, Netto-Ferreira, André L., Ota, Rafaela P., T. Oyakawa, Osvaldo, Pavanelli, Carla S., Reis, Roberto E., Santos, Osmar, Serra, Jane Piton, Silva, Gabriel S. C., Silva-Oliveira, Cárlison, Souza-Lima, Rosana, Vari, Richard P. & Zanata, Angela M. 2024 |
Nannocharax reidi
Vari RP & Ferraris CJ, Jr. 2004: 552 |
Nannocharax rubrolabiatus
Van den Bergh EC & Teugels GG & Coenen EJ & Ollevier F. 1995: 350 |
Neolebias lozii
Winemiller KO & Kelso-Winemiller LC 1993: 112 |
Nannocharax maculicauda Vari, Géry, 1981:1076
Vari RP & Gery J. 1981: 1076 |
Onouphrios
Whitley GP 1951: 68 |
Distichodus tanganicae
Worthington EB & Ricardo CK 1937: 1083 |
Nannocharax minutus
Worthington EB 1933: 40 |
Nannocharax niloticus var. lateralis
Werner F. 1906: 1130 |