Pantoclis Förster, 1856

Hübner, Jeremy, Chemyreva, Vasilisa, Macek, Jan & Kolyada, Victor, 2024, A review of the genus Zygota (Hymenoptera, Diapriidae) in Germany with taxonomic notes on this genus and its distinction from Pantoclis, ZooKeys 1207, pp. 325-353 : 325-353

Genus Pantoclis Förster, 1856

Type species.

Pantoclis barycera Förster, 1861 (Figs 1 A, B , 5 E ).

Diagnosis.

Body black to yellowish brown; males macropterous, females alate to brachypterous or wingless; occipital carina always with occipital pit (Fig. 1 B , red arrow); fore tibiae of males always unmodified with homogeneous pubescence (Fig. 3 H ); submetapleural carina usually present, complete (Fig. 2 A , green arrow) [if submetapleural carina missing, then venation as described below]; radial cell open to closed, variable in shape (Fig. 14 ); radialis not parallel to parastigma [if parallel (Fig. 3 G, J ) then angle between stigmal and marginal veins as described below]; angle between stigmal and marginal veins 130 degrees (Fig. 3 G, J ) or more; S 2 always smooth, without punctured area on it in anterior half (Fig. 4 F ); male genitalia usually slender, apex of aedeagus distinctly convex (Fig. 5 I – L ), lanceolate (Fig. 5 F – L ), rather truncate (Fig. 5 E ) [if genitalia short and stout with rounded aedeagus then fore wing with a closed radial cell], digitus usually diminished (Fig. 5 E – L ) [if not then fore wing with closed radial cell]; ovipositor usually long, at least as long as length of T 2 [if ovipositor short then fore wing with closed radial cell].

Figures

Figure 1. Morphological characters to identify the closely related genera Zygota and Pantoclis A, E female B, C, D males A, B P. barycera C Z. walli sp. nov. D, E Z. abdominalis. Scale bars: 1 mm (A); 0.5 mm (B – F).

Figure 2. Morphological characters to identify the closely related genera Pantoclis (A) and Zygota (B) A Pantoclis spp., male B Z. breviuscula, male. Green arrow – submetapleural carina. Scale bars: 0.3 mm.

Figure 3. Venation (A, C, D, G, J) and fore tibia (B, E, F, H, I) morphology of males A Zygota bensoni B Z. sordida C Z. croton D Z. walli sp. nov. E Z. breviuscula F Z. walli sp. nov. H Pantoclis sp. I Z. croton G, J Pantoclis spp. Scale bars: 0.5 mm (A – E, G, I, J); 0.3 mm (F, H).

Figure 4. Ventral side of metasoma of females (A, C, E) and males (B, D, F) A Zygota breviuscula B Z. abdominalis C, D Z. pubescence E Z. walli sp. nov. F Pantoclis sp. Scale bar: 0.5 mm.

Figure 5. Male genitalia of Zygota and Pantoclis A Z. walli sp. nov. B – D Z. abdominalis E P. barycera F – H Pantoclis sp. 1 I – L Pantoclis sp. 2 C, G, I, K lateral view A, B, D, E, F, H, J, L ventral view.

Figure 14. Phylogenetic ML tree of barcoded Pantoclis material and the polyphyletic appearance of their wing venation. Green represents the taxa with a closed radial cell while species with an open cell are color-coded red. Each node’s support is displayed by the bootstrap and the jackknife values. Aclista was used as an outgroup.