Pupina verneaui Dautzenberg & Fischer, 1906

Jirapatrasilp, Parin, Sutcharit, Chirasak & Panha, Somsak, 2022, Annotated checklist of the operculated land snails from Thailand (Mollusca, Gastropoda, Caenogastropoda): the family Pupinidae, with descriptions of several new species and subspecies, and notes on classification of Pupina Vignard, 1829 and Pupinella Gray, 1850 from mainland Southeast Asia, ZooKeys 1119, pp. 1-115 : 1

Pupina verneaui Dautzenberg & Fischer, 1906

Figs 21T , 26D

Pupina verneaui Dautzenberg & Fischer, 1906 [1905]: 440, 441, pl. 10, figs 13-15. Type locality: Ha-Giang [Ha Giang Province, Vietnam]. Fischer 1963: 34. Do et al. 2015: 126, fig. 6c, Son La Province, Vietnam.

Eupupina verneaui - Dautzenberg and Fischer 1908: 208, 209, Mo-Xat [west of Quang Uyen, Cao Bang Province, Vietnam]; Quang-Huyen [Quang Uyen, Cao Bang Province, Vietnam].

Type material examined.

Syntypes MNHN-IM-2000-35843 (Figs 21T , 26D ) from Ha-Giang, Tonkin.

Diagnosis.

Shell ovate-fusiform; last whorl ca. 70% of shell height; suture very shallow. Apertural lip somewhat thickened, not expanded. Both parietal and columellar teeth fin-shaped and thickened; parietal tooth somewhat covering posterior canal; columellar tooth next to slit-like anterior canal.

Differential diagnosis.

Pupina verneaui is most similar to P. artata in having fin-shaped and thickened teeth, but differs in having a more ovate-fusiform shell shape and a rather shallower suture.

Distribution.

Northern Vietnam ( Do et al. 2015).

Remarks.

The specimen of P. verneaui figured in Inkhavilay et al. (2019: fig. 16a) from Ban Nong Kham village, Kasy District, Vientiane Province, Laos should constitute a different species as it is different from the syntype figured here in having a wider spire, a more bulging last whorl and a thinner and sharper parietal tooth.

Figures

Figure 21. Shells of Pupina artata species group from mainland Southeast Asia A-M Pupina artata A syntype UMZC I. 102960. A and specimens B CUMZ 12011 C CUMZ 12023 D CUMZ 12038 E CUMZ 12026 F CUMZ 12015 G CUMZ 12009 H CUMZ 12001 I CUMZ 12003 J NHMUK 1906.4.4.28 K CUMZ 12037 L CUMZ 12031, and M CUMZ 12018 N, O Pupina pallens N lectotype SMF 109951 and O specimen CUMZ 12042 P-R Pupina limitanea P, Q syntypes NHMUK 1903.7.1.2967 and R specimen CUMZ 12044 S Pupina billeti, holotype MNHN-IM- 2000 - 35841 T Pupina verneaui, syntype MNHN-IM- 2000 - 35843 U, V Pupina hungerfordiana, specimens NHMUK 91.3.14.686 - 7 W, X Pupina bensoni sp. nov. W holotype CUMZ 12045 / 1 and X paratype CUMZ 12046 / 1. Photo: H. Taylor, NHM (A, P, Q), P. Maestrati, MNHN (S, T).

Figure 26. A Pupina bensoni sp. nov., paratype CUMZ 12046 / 1 from Khao Wong Cave, Uthai Thani B Pupina hungerfordiana, specimen NHMUK 91.3.14.686 - 7 from Hsaddan Koo C Pupina billeti, holotype MNHN-IM- 2000 - 35841 D Pupina verneaui, syntype MNHN-IM- 2000 - 35843 from Ha-Giang, Tonkin E, F Pupina peguensis E syntype of Pupina blanfordi NHMUK 1888.12.4.100 from Pegu and F specimen NHMUK ex. Cuming coll. from Lao Mountains, Camboja. Photo: P. Maestrati, MNHN (C, D).