Erythromelana Townsend, 1919

Fleming, AJ, Wood, D. Monty, Smith, M. Alex, Hallwachs, Winnie, Janzen, Daniel & Dapkey, Tanya, 2016, Two new species of Erythromelana Townsend, 1919 (Diptera: Tachinidae) from Area de Conservacion Guanacaste in northwestern Costa Rica, Biodiversity Data Journal 4, pp. 7386-7386 : 7386

Erythromelana Townsend, 1919

Erythromelana Erythromelana Townsend, 1919

Erythromelana Townsend, 1919a: 174. Type species: Erythromelana jaena Townsend, 1919a, by original designation. Other references: Guimarães (1971: 106); Wood (1985: 39-40); Wood & Zumbado (2011: 1403).

Minthomyia Townsend, 1919b: 564. Type species: Minthomyia abdominalis Townsend, 1919b, by original designation. Other references: Guimarães (1971: 41); Wood (1985: 39-40) (as synonym of Erythromelana ).

Erythromelana Erythromelana jaena Townsend, 1919 Townsend 1919: 174.

Description

Erythromelana can be distinguished from other Blondeliiniby the following combination of characters: Head: proclinate orbital setae absent in male, female with 2 setae; 1-2 pairs of reclinate outer orbital setae, lowermost outer orbital seta distinctly longer than uppermost frontal seta; ocellar setae ranging from absent to well developed; eyes haired, of variable density in different species; parafacial bare and extremely narrow, at narrowest point equal to or narrower than the basal width of the palpus; parafacial light gray in ground color, covered with silvery–gold pollinosity, and bare; fronto-orbital plate and vertex black in ground color, covered with a dull silver-gold pollinosity (appearing mostly black (Fig. 2b)), and with faint golden reflections visible only in lateral view (mostly on vertex); lower margin of face level with vibrissa; vibrissa positioned at extreme anteroventral corner of face; facial ridge with a few short hairs on lower third or less; arista black on basal 1/3-1/4, becoming light brownish to orange distally, elongate, and minutely pubescent, thickened only on basal 1/4; palpus light orange to yellow, sometimes darkened basally. Thorax: ground color shiny black; scutum silver pollinose presuturally, postsuturally a polished black in ground color; prosternum setose; postpronotum bearing 2 or, rarely, 3 setae; katepisternum with 2-3 setae; first postsutural supra-alar seta small or sometimes absent; apical scutellar setae lacking; wingvein R4+5 setulose at base and vein R1 setulose or bare; vein M smoothly curved at bend and ending at wing margin anterior to wing tip, separately from vein R4+5; legs ranging from entirely yellow to entirely black. Abdomen: ground color ranging from yellow-orange to black, without strong banding; mid-dorsal depression not extending to hind margin of T1+2; discal setae only present on T5. Male terminalia: sternite 5 with median cleft either U- or V-shaped; inner margin bearing minute setae; apical lobe rounded or pointed apically with either a single, long, well-developed seta, multiple well developed setae, or bare; pregonite curved anteriorly, with strong setae along posterior margin; postgonite short with curved apex; epiphallus small, usually difficult to see between the pregonites; surstylus with setulae on inner and outer surfaces, or completely bare; surstylus, in lateral view, varied in shape from almost straight to slightly concave on anterior or posterior margins, usually with a broad, rounded apex, occasionally truncate; surstylus and cercus usually subequal in length, sometimes cercus shorter than surstylus; in posterior view, cerci narrowed on apical 1/3.

Other species included

Based on synapomorphieswithin the male terminalia of Erythromelana described by Inclan and Stireman (2013), the two newly described species belong to the E. cryptica species-group and as such were compared to the other species belonging to that group.

arciforceps Inclan, 2013: 35. Holotype male (CNC). Type locality: Brazil, S.C., Nova Teutonia. Type label: Nova Teutonia S. C.-Brazil Nov. 1970 F. Plaumann/ HOLOTYPE Erythromelana arciforceps Inclan D.J/ DI412CA. Additional specimens examined: Brazil. [Examined by AJF.]

catarina Inclan, 2013: 32. Holotype male (CNC). Type locality: Brazil, S.C., Nova Teutonia. Type label: Nova Teutonia S. C.-Brazil Jun. 1970 F. Plaumann/ HOLOTYPE Erythromelana catarina Inclan D.J/ DI392CA. Additional specimens examined: Brazil. [Examined by AJF.]

convexiforceps Inclan, 2013: 33. Holotype male (CNC). Type locality: Mexico, Oaxaca, Suchistepec. Type label: Mexico, Oax 4.6 km Suchistepec 23.VII.1992 D.M. Wood 2150m/ HOLOTYPE Erythromelana convexiforceps Inclan D.J./ DI54CA. [Examined by AJF.]

cryptica Inclan, 2013: 29. Holotype male (CNC). Type locality: Venezuela, Aragua, Rancho Grande. Type label: VENEZUELA Aragua Rancho Grande 18-27.II.1971 G.&M. Wood 1100m/ HOLOTYPE Erythromelana cryptica Inclan D.J./ DI477CA. Additional specimens examined: Venezuela, Bolivia, Mexico and Ecuador. [Examined by AJF.]

distincta Inclan, 2013: 39. Holotype male (CNC). Type locality: Venezuela, Aragua, Rancho Grande. Type label: VENEZUELA Aragua 11 km Rancho Grande 25.II.1971 G.&M. Wood/ HOLOTYPE Erythromelana distincta Inclan D.J./ DI280CA. Additional specimens examined: Brazil and Venezuela. [Examined by AJF.]

napensis Inclan, 2013: 37. Holotype male (CNC). Type locality: Ecuador, Napo prov., Yanayacu Biological Station. Type label: ECUADOR: Napo Prov. Yanayacu Biological Station S00°35.9', W77°53.4', 2163 m REARED October 2005 8135/ HOLOTYPE Erythromelana napensis Inclan D.J. [Examined by AJF.]

woodi Inclan, 2013: 42. Holotype male (CNC). Type locality: Costa Rica, Puntarenas, Monteverde. Type label: COSTA RICA Pnts Monteverde 28.VIII.1993 D.M. Wood 1842m/ HOLOTYPE Erythromelana woodi Inclan D.J./ DI208MW. Additional specimens examined: Costa Rica, Bolivia, Mexico and Ecuador. [Examined by AJF.]