Hurkaea conringioides subsp. conringioides

Koch, Marcus A., Karl, Robert, Al-Shehbaz, Ihsan A. & German, Dmitry A., 2022, The new tribe Fourraeeae (Brassicaceae) and two Moroccan Arabis species transferred to the new genus Hurkaea, Phytotaxa 543 (1), pp. 41-50 : 46-47

Hurkaea conringioides subsp. conringioides

= Arabis werneri Emb. & Maire ( Maire 1933: 196) . ≡ A. conringioides var. werneri (Emb. & Maire) Galland & Favarger (1985: 233) . Type indication: “Hab. in fissuris rupium vulcanicarum Anti-Atlantis: in ditionis Siroua monte Amezdour, ad alt. 2600–2700 m ”. Lectotype (designated here):— MOROCCO: Siroua, Au Tizi-n-Ougdar, Mt. Amezdour (as Amerouz), 2600 m, 30 June 1932, L. Emberger s.n. (MPU-002817; isolectotype: MPU-002818).

Hurkaea conringioides subsp. humbertii (Quézel) Al-Shehbaz, M. Koch & R. Karl , comb. nov. ≡ Arabis humbertii Quézel (1953: 198) . ≡ Arabis conringioides subsp. humbertii (Quézel) Dobignard & Jordan (1987: 79) . Type indication: “Hab. in atlante majore orientale, loco vocato Imi N’Tamd.” Type:— MOROCCO: J. Ayachi, Imi-N-Tamd, June-July 1952, C. Guinet, Ch. Souvage et R. de Villmorin 792 (RAB-000465).

Note. Galland & Favarger (1985) recognized three varieties in Arabis conringioides , including var. conringioides, var. werneri , and var. aphanostyla Maire. The last was invalidly published in Maire (1967). By contrast, Dobignard & Jordan (1987) recognized the two subspecies listed here, though they accepted var. aphanostyla , which Dobignard et al. (2011) subsequently reduced to synonymy of subsp. humbertii .

Hurkaea josiae (Jahand. & Maire) Al-Shehbaz, M. Koch, R. Karl, & D.A. German , comb. nov. ≡ Arabis josiae Jahandiez & Maire (1923: 66) . Type indication: “Hab. in silvis (quercetis et cedretis) Atlantis Medii maroccani supra oppidum Azrou, ubi maio floret – Typus in Herbario Universitatis Algeriensis. … Cette plante a été ensuite récoltée en fleurs, le 16 mai 1921, per Jahandiez, dans le cratère du Djebel Hebbri, entre Azrou et Timhadit, dans les clairières des cédraires, sur basalte vers 1900 m. Elle a été ensuite retrouvée par Maire dans les forêts de Quercus ilex au dessus d’Azrou, sur calcaire jurassique vers 1400 m, en fruits plus que mûrs, le 25 juillet 1921 ”. Type: Morocco, Moyen Atlas , above town Azrou, 1921, É. Jahandiez &/or R. Maire s.n.

Note. Although the North African herbarium of the University of Algeria (AL) is listed in Index Herbariorum online (sweetgum.nybg.org/science/ih/), no address or correspondents are given, and inquiry about the type collection of Arabis josiae was not possible. The original protologue ( Jahandiez & Maire 1923) lists two gatherings, and relevant syntypes are available in MPU (with printed labels from AL “Université d’Algiere Herbier del’Afrique du Nord”) and AIX. The collections by Jahandiez were made on 16 May 1921 (MPU-000159, AIX-000066) at 1900 m from Cedrus forests, and they are immature plants in early stages of flowering. However, Maire’s collections (MPU-000157, MPU-000158) were made on 25 July 1921 at 1400 m from Quercus forests. They are with fully mature plants. Because the original description includes flowering and fruiting material, and because the fruits are essential for the characterization of the genus Hurkaea , a designation of the specimen with mature fruits at AL as the lectotype would be preferable, and its duplicate at MPU would then be the isolectotype.

= Arabis leptopoda Pau & Font Quer ( Font Quer 1928: no 236, in sched.). ≡ A. josiae var. leptopoda (Pau & Font Quer) Pau & Font Quer ( Font Quer 1929: no 139, in sched.). Type indication: “Hab. in saxosis, lacisque silvaticis montis Iguermalez (Atlante rhiphaeo), 1950 m. alt.; 7 junii”. Lectotype (designated by Nualart et al. 2021: 14):— MOROCCO: In saxosis, lacisque silvaticis montis Iguermalez (Atlante rhiphaeo), 1950 m, 7 June 1928, Font Quer 236-1928 (MA-48513; isolectotypes BC-2978, BCN-17301, BM-000583666, GDA-28946, MPU-006331, MPU 006332).