Fourraea Greuter & Burdet ( Greuter & Raus 1984: 283 )

Koch, Marcus A., Karl, Robert, Al-Shehbaz, Ihsan A. & German, Dmitry A., 2022, The new tribe Fourraeeae (Brassicaceae) and two Moroccan Arabis species transferred to the new genus Hurkaea, Phytotaxa 543 (1), pp. 41-50 : 45

Fourraea Greuter & Burdet ( Greuter & Raus 1984: 283)

Type: Fourraea alpina (L.) Greuter & Burdet.

= “ Caulopsis ” Fourreau (1868: 332), nom. inval.

Herbs perennial, with elongated caudex, glaucous. Trichomes simple, confined only to petioles. Multicellular glands absent. Stems erect, simple, leafy, unarmed. Basal leaves long petiolate, ciliate along proximal half of petiole, rosulate; blade simple, entire, sometimes ciliate near base; cauline leaves sessile, amplexicaul to strongly auriculate at base, simple, entire. Racemes few to many flowered, ebracteate throughout, lax, elongated considerably in fruit; rachis straight; fruiting pedicels ascending-divaricate, slightly curved or straight, persistent. Sepals elliptic-oblong, free, deciduous, erect, equal, base of lateral pair not saccate. Petals white, erect to ascending, longer than sepals; blade narrowly oblanceolate, apex rounded; claw shorter than sepals, glabrous, unappendaged, entire. Stamens 6, slightly exserted, erect, tetradynamous; filaments wingless, unappendaged, glabrous, free; anthers oblong, apex obtuse. Nectar glands 4, distinct or confluent; median nectaries ovoid, smaller than laterals; lateral nectaries ringlike. Ovules (30–)36–52 per ovary; placentation parietal. Fruit dehiscent, capsular siliques, linear, erect to ascending, subtetragonal-latiseptate, not inflated, unsegmented; valves thin or thick leathery, with a prominent midvein, glabrous, not keeled, slightly torulose, wingless, unappendaged; gynophore obsolete or to 0.5 mm long; replum rounded, visible; septum complete, membranous, veinless; style absent, obsolete, or 0.7–1 mm long, persistent; stigma capitate, entire, unappendaged. Seeds uniseriate, wingless, oblong-ovoid, plump; seed coat mucilaginous when wetted; cotyledons obliquely accumbent.

A monospecific European genus.

Fourraea alpina (L.) Greuter & Burdet ( Greuter & Raus 1984: 283). ≡ Brassica alpina Linnaeus (1767: 95) .

For a complete synonymy, see BrassiBase (https://brassibase.cos.uni-heidelberg.de), POWO (https://powo. science.kew.org/), WCVP (https://wcvp.science.kew.org/) [the latter two under Turritis brassica Leers (1775: 147) ], and for species descriptions and distributions see Akeroyd (1993) and Jalas & Suominen (1994), respectively, both under Arabis brassica (Leers) Rauschert (1973: 648) , as well as Talavera (1993) under A. pauciflora and Euro+Med PlantBase (https://europlusmed.org/).

Distribution: Andorra, Austria, Belgium, Croatia, Czech Republic, France, Germany, Italy, Liechtenstein, Luxembourg, Serbia, Slovakia, Slovenia, Spain, Switzerland.