Philmontoides globosus, Ingrisch, 2022

Ingrisch, Sigfrid, 2022, Revision of the genus Philmontis Willemse, 1966 and description of a new genus Philmontoides gen. nov. from New Guinea (Orthoptera: Tettigoniidae; Conocephalinae), Zootaxa 5182 (2), pp. 101-151 : 135-137

publication ID	https://doi.org/ 10.11646/zootaxa.5182.2.1
publication LSID	lsid:zoobank.org:pub:8920DE84-2BE6-4A68-A7F7-AC987F1F894E
DOI	https://doi.org/10.5281/zenodo.7053843
persistent identifier	https://treatment.plazi.org/id/DB181868-FFBA-FFE9-FF67-D52329E5F17E
treatment provided by	Plazi
scientific name	Philmontoides globosus
status	sp. nov.

Treatment

Philmontoides globosus sp. nov.

Figs. 9G–H View FIGURE 9 , 10E–F View FIGURE 10 , 11C View FIGURE 11 , 12C View FIGURE 12 , 14D–G View FIGURE 14 , 16I–L View FIGURE 16

Holotype (male): Indonesia: Papua, S. Geelvink Bay, Nabire , elev. 3 m (3°22’S, 135°29’E), 3.xii.1962, leg. J.L. Gressitt —depository: Bernice B. Bishop Museum, Honolulu ( BPBM). GoogleMaps

Other specimens studied: Indonesia: same data as holotype— 2 males ( BPBM); Papua [New Guinea NW], Nabire, Geelvink Bay ( Cenderawasih Bay ), elev. 0–30 m (3°22’S 135°30’E), 2–9.vii.1962, leg. J. Sedlacek — 1 female ( BPBM) GoogleMaps .

Diagnosis. Males of the new species have the general appearance and the abdominal appendages similar to those of P. lobatus ( Naskrecki & Rentz, 2010) , but the male cerci are less strongly curved in P. globosus sp. nov. than in P. lobatus and have the apical margin of the widened ventral apical branch little concave instead of angularly excised. The titillators of both species have a similar basic plan, but in P. globosus the proximal apical branches are wider and longer and semi-oval instead of wide triangular, and the distal branches shorter and more strongly vaulted than in P. lobatus . The female subgenital plate of P. globosus is similar to or identical with that of P. lobatus although in the specimens at hand it appears wider and shorter in P. globosus while narrower and longer in P. lobatus . But the differences might be due to distortions or individual variation. Thus, females of both species cannot be differentiated with certainty without corresponding males. The female subgenital plate of P. globosus is also similar to that of P. hageni ( Dohrn, 1905) . It differs by the baso-lateral extensions, which in P. globosus arise from the latero-anterior angles of the plate and are thus fixed to the plate while in P. hageni they arise from the very base of the plate. Males of P. globosus differ from those of P. hageni by the narrow, about cylindrical male cerci instead of being widened with concave internal surface and by the shape of the titillators.

Description. Elongate species with tegmen nearly reaching end of hind femur ( Figs 9G–H View FIGURE 9 ). Pronotum elongate; disc with anterior margin convex but faintly concave in middle, posterior margin broadly rounded, covering only very base of stridulatory area of tegmen in male; lateral lobes not very deep, ventral margin slightly concave, at end forming a rounded angle with posterior area that is projecting laterad and clearly visible from above. Prosternal spines long; mesosternal lobes obtuse; metasternal lobes rounded with an angle or fold. Femora with the following number of spines on ventral margins: (1) 3-8 / 6-8; (2) 6-9 / 1-4; (3) 11-13 / 9-15 (n = 4); hind knee lobes bispinose; in one male also anterior knee lobe of fore femur bispinose.

Male. Stridulatory file in about basal two thirds with large and spaced teeth, after a sinusoidal step, teeth abruptly narrowing and becoming denser and hardly perceptible toward end; total file length 1.32 mm with 38 distinct and about 10 indistinct teeth; area with countable teeth 1.29 mm with 38 teeth; area with large teeth before step 0.92 mm with 17 teeth; area with largest teeth 0.68 mm with 12 teeth (n=1; Fig. 12C View FIGURE 12 ). Tenth abdominal tergite strongly convex with short rounded excision from base, in apical area markedly down curved, hind margin on both sides concave, in middle prolonged into a process with concave lateral margins that shortly after divides into a pair of compressed projections that widen toward tip and embrace an oval space that is open at hind margin ( Fig. 11C View FIGURE 11 ). Cerci with basal rim forming a narrow furrow with a small bump at both sides, afterward rounded, slightly conical and little curved mediad, terminating into a dorsal and a ventral projection; dorsal projection compressed, with narrowing margins, at end rounded with a small, curved spine; ventral projection wide, dorso-ventrally compressed, at end with a triangular ventral projection with straight hind margin and with a small spine below its dorsal hind angle ( Figs 10E–F View FIGURE 10 ). Subgenital plate strongly, triangularly excised from base to almost mid-length, afterward with a fine carinula along midline; central disc at base slightly convex, afterward with little converging lateral margins; margins upcurved and at dorsal end curved laterad, forming in basal half a narrow concave, in apical half a wide convex rim; the plate terminates into a pair of narrow, about rounded projections that carry styli at oblique ventroapical end ( Fig. 11C View FIGURE 11 ). Titillators with basal area long and narrow, running parallel but not fused with each other, at base for a short distance curved sideward; at posterior end curved moderately sideward and widened to vaulted laminar branches; these branches appear in apical view to have the dorsal rims curved in an about 180° angle against each other while the lateral surfaces are strongly bulging on both sides with internal surface more strongly bulging and shorter, external surface less strongly bulging and longer ( Figs 14D–G View FIGURE 14 ). There is also a pair of narrow, wavy lateral sclerites near bases of titillators running vertically to the titillator axis.

Female. Subgenital plate with baso-lateral expansions large, oval, and moderately swollen; disc little bent along mid-line forming a stout medial carina; lateral margins converging, ending into a pair of triangular extensions with obtuse tips, hind margin angularly excised in between ( Figs 16I–L View FIGURE 16 ).

Coloration. Face of general color, antennal scrobae light or darkened, internal surface of scapus and fastigium verticis brown or black; vertex, occiput and upper area of genae black or dark brown; dorsal surface of pronotum of general color, in posterior area with indication of a dark medial band, lateral lobes black with one or two light spots, margins black but partly of light color; tegmina along anterior and posterior margins of light color, partly with dark spots, in central area dark brown with light veinlets, along central area thickened and blackish brown; hind femur with black pre-genicular ring ( Fig. 16I View FIGURE 16 ).

Measurements (3 males, 1 female).—Body w/wings: male 32.5–35.0, female 36; body w/o wings: male 26–30, female 33; pronotum: male 6.5–7.8, female 7; tegmen: male 22–25, female 26; hind femur: male 21.5–23.0, female 24; antenna: male 120–130; ovipositor: female 14 mm.

Etymology. The new species is named for the globular appearance of the male titillators in apical view; from Latin globosus globular.