Tachyusa concinna Heer, 1839

Pa ṡnik, Grzegorz, 2006, A revision of the World species of the genus Tachyusa Erichson, 1837 (Coleoptera, Staphylinidae: Aleocharinae), Zootaxa 1146, pp. 1-152 : 119-124

Tachyusa concinna Heer, 1839 ( Figs. 123–125)

Tachyusa concinna Heer, 1839: 345 .

Tachyusa concinna: Mulsant & Rey, 1875: 390 (as valid species). Tachyusa concinna: Fauvel, 1895: 98 (as synonym of T. coarctata ).

Tachyusa (s. str.) concinna: Reitter, 1909: 73 (as synonym of T. coarctata ).

Tachyusa (s. str.) concinna: Bernhauer & Scheerpeltz, 1926: 584 (as synonym of T. coarctata ).

Type material. Neotype: ♂: Switzerland: Aarau ( NMW).

Additional material. Austria: Albern , Donau, Scheerpeltz, 1 ex ( NMW) ; Altenburg, P. Heymes, 1 ex ( ZMHB) ; Drau River bank near Ferlach , 1–12.VIII.1924, Scheerpeltz, 3 exx ( NMW) ; Dresden , 1 ex ( FMNH) ; Eisenkappel, Karnten, Sandbank Miklautzhof , 7.VIII.1931, Scheerpeltz, 1 ex ( MHNG) ; Gollingen , 22.VI.1953, 2 exx ( ZMHB) ; Haslau, Scheerpeltz , 1 ex ( MHNG) ; Inzeradorf bei Wien, Liestngbach river , 28.V.1946, P. Meyer, 1 ex ( NMW) ; Kampthal, Bernhauer , 1 ex ( FMNH) ; 2 exx ( MHNG); 3 exx ( NMW); 1 ex ( MTD); Lobau near Wien, Scheerpeltz, 2 exx ( NMW) ; Marchfeld, Baumgarten , 6.VII.1965, 6 exx ( ZMHB) ; Stockerau, Bernhauer , 1 ex ( FMNH) ; 1 ex ( ZMHB); 1 ex ( NMW); Weidlingbch, Breit , 1 ex ( NMW) ; Wien, 2.VI.1946, Meyer, 1 ex ( MHNG) ; Wien, 4 exx ( NMW) ; Wien, Tullnerbach , 2 exx ( FMNH) ; 1 ex ( MTD); Wien, Moczarski , 1 ex ( NMW) ; “ Donauauen ” (localities not specified), 1 ex ( MCZ) ; 1 ex ( NMW). Belarus: Pińsk , VI.1938, Mazur, 1 ex ( MIZ) ; Siennica, powiat Minski , 22.IV.1934, 3 exx ( MIZ) . Belgium: Anderlecht, Pede valley , 9.V.1945, Fagel, 1 ex ( IRSNB) ; Anvers , 18.VI.1938, Fagel, 4 exx ( IRSNB) ; 23.V.1948, Fagel, 1 ex ( IRSNB) ; Auderghem, Rouge­Cloitre , 19.VII.1945, Fagel, 1 ex ( IRSNB) ; Olloy, Viroin , 23.VI.1953, Fagel, 1 ex ( IRSNB) ; Ruysbroeck , 17.IV.1943, Fagel, 2 exx ( IRSNB) ; Seilles, Sclaigneaux , 17.VIII.1945, Fagel, 4 exx ( IRSNB) ; Uccle, Val d'Engeland , 12.VII.1945, Fagel, 1 ex ( IRSNB) ; 3.IX.1945, Fagel, 1 ex ( IRSNB) ; Wemmel , 17.V.1936, 1 ex ( IRSNB) . Bosnia and Herzegovina: Bjelasnica, 1 ex ( IRSNB) ; Ilidža , 1 ex ( MTD) . Bulgaria: Sandanski , 16–23.VII.1985, M. Schülke, 4 exx ( ZMHB) ; Strumaufer near Kresna , 29.V.1984, H. Wendt, 1 ex ( VAPC) . Czech Republic: Bechovice , 6.VII.1907, Dr. Uzel, 4 exx ( NMPC) ; “ Brandeis ” [Brandys], Skalitzky, 2 exx ( NMW) ; Čelakovice , VI.1906, Rambousek, 6 exx ( NMPC) ; 10.VI.1906, Klička, 5 exx ( NMPC) ; 13.VIII.1911, Dr. Klička, 1 ex ( NMPC) ; Chuchel, Lokay , 1 ex ( NMPC) ; Davle, Svoboda , 2 exx ( NMPC) ; “ Jungbunzlau ” [Mlada Boleslav], Skalitzky, 1 ex ( NMW) ; Libic , 26.V.1907, Klička, 4 exx ( NMPC) ; Lobkovice, Zeman , XI.1905, 16 exx ( NMPC) ; Mut nice, J. Bechyne, 5 exx ( NMPC) ; Nymburk, Zeman , 2 exx ( NMPC) ; Pardubice, Zeman , 1 ex ( NMPC) ; “ Paskau ” [Paskov], Reitter, 4 exx ( NMW) ; 1 ex ( SMNS); Praha Rubny , 17.VII.1907, 1 ex ( NMPC) ; Praha, V.M. Duchon, 7 exx ( NMPC) ; Praha, 18.VI.1929, 6 exx ( NMPC) ; “Prossnitz" [ Prostejov ], V. Zoufal, 1 ex ( ZMHB) ; 18 exx ( NMPC); Roudnice , V.1907, 1 ex ( NMPC) ; 28.V.1907, Roubal, 1 ex ( DEI) ; Vrané, Smolka , 1 ex ( MIZ) ; “ Wilken ” [Vlkan], Hildesheim, 2 exx ( NMPC) ; Wran near Praha, Dr Rodt, 1 ex ( NMPC) ; Zlichov , 2.VI.1901, 1 ex ( NMPC) ; 11.VI.1905, Rambousek, 1 ex ( NMPC) . Estonia: “ Dorpat ” [Tartu], 21.VIII.1896, Klefbeck, 1 ex ( MZLU) ; Pärnu, 25.VII–4.VIII.1938, Palm, 8 exx ( MZLU) . Finland: Pyhäjärvi , Sahlberg, 1 ex ( NMW) . France: Allier , 1 ex ( MNHN) ; Ancenis , 1 ex ( MNHN) ; Anjou , 1 ex ( MNHN) ; Bonneuil , VIII.1938, Levasseur, 1 ex ( MNHN) ; Boulogne, Seine River , 11 exx ( MNHN) ; Briare Loizet , 17.I.1943, Fongond, 2 exx ( MNHN) ; Chatou , V.1935, 2 exx ( MNHN) ; V.1936, 9 exx ( MNHN) ; V.1939, 5 exx ( MNHN) ; IV.1942, 3 exx ( MNHN) ; Fort­Louis, Rhin , IX.1929, 4 exx ( MNHN) ; Gennevilliers , 16.V.1924, 5 exx ( MNHN) ; 16.VIII.1924, 4 exx ( MNHN) ; Lille , 3 exx ( BMNH) ; Mello, Oise , 1 ex ( MNHN) ; Nevers , 1903, 1 ex ( MNHN) ; Paris , 7 exx ( BMNH) ; Paris , 3 exx ( MNHN) ; Semur, Cote d'Or , IX.1942, 1 ex ( MNHN) ; Soissons , 4 exx ( MNHN) ; Saint Dizier, Marne , Deville, 7 exx ( MNHN) ; Saint Florent, Cher , Deville, 1 ex ( MNHN) ; Strasbourg , IX.1953, Coiffait, 26 exx ( MNHN) ; Terraube, H. Belliard, 1 ex ( NMW) ; Villeneuve , V.1942, 1 ex ( MNHN) ; VI.1947, Levasseur, 2 exx ( MNHN) ; Vitry , 3.VIII.1941, 1 ex ( MNHN) ; VII.1961, 1 ex ( MNHN) ; Wissous , V.1889, 1 ex ( MNHN) ; Normandie (exact locality not specified), 2 exx ( NMW). Georgia: Tbilisi, Mzcheta , 16.VII.1985, Wrase, 1 ex ( ZMHB) ; Tbilisi citi border, 6.VII.1987, Schülke & Wrase, 1 ex ( ZMHB) . Germany: 1.5 km W Münchehofe, Elsengrund Bachufer , 25.V.1985, M. Uhlig, 1 ex ( ZMHB) ; Alt Ruppin, Tornowsee River , 8.VII.1922, 1 ex ( ZMHB) ; 10.VII.1922, 1 ex ( ZMHB) ; Altenberg, Lokay , 1 ex ( NMPC) ; Arnstadt, Thur. , 6.VI.1950, W. Liebmann, 1 ex ( DEI) ; Berlin, K. Schubert, 3 exx ( CAS) ; 26 exx ( ZMHB); Wagner , 4 exx ( DEI) ; 30.VII.1904, 2 exx ( CAS) ; 28.V.1907, Roubal, 4 exx ( DEI) ; 3 exx ( DEI); 3 exx ( ZMHB); Boizenburg , 2 exx ( MHNG) ; Brieselang near Nauen , 17.V.1917, 2 exx ( ZMHB) ; Brodowin , 14.VII.1928, 2 exx ( ZMHB) ; Chemnitz, Kirsch , 2 exx ( MTD) ; Corenick near Berlin, Wendeler, 1 ex ( ZMHB) ; Darss Prerow, MTB 1540, Weststrand , 29.VII.1986, Wrase, 1 ex ( ISEA) ; 6 exx ( ZMHB); Dessau, E. Heidenreich, 5 exx ( MHNG) ; 1 ex ( MTD); IV.1909, 4 exx ( ZMHB) ; Dossow , 1918, K. Hanel, 1 ex ( MTD) ; Dresden , 15.V.1908, Blasewitz, 1 ex ( MTD) ; Kirsch , 4 exx ( MTD) ; Ebersbach, Lobau , 20.VI.1973, Hieke, 1 ex ( ZMHB) ; Elienburg , 9.VII.1958, Linke, 4 exx ( MTD) ; Erlangen, Bauer , 1 ex ( DEI) ; Finkenkrug , 3 exx ( DEI) ; 1918, K. Hanel, 10 exx ( MTD) ; Garmisch , 18.VII.1934, Ihssen, 1 ex ( ZMHB) ; Geesthacht near Hamburg, 22.V.1893, Koltze, 8 exx ( DEI) ; 9.VII.1893, 6 exx ( NMPC) ; Gorlitz , 26.V.1977, Vogel, 1 ex ( AZPC) ; 12.VI.1976, Vogel, 2 exx ( ZMHB) ; Grunbusch near Hirschberg , Maischner, 1 ex ( ZMHB) ; Guttau, Kreis Bautzen , 28.V.1982, M. Schülke, 1 ex ( ZMHB) ; Halle , 7 ex ( ZMUC) ; Kaiserstuhl , 5.VI.1943, 1 ex ( ZMHB) ; Kladow am Wannsee , 1 ex ( MIZ) ; Kleinheringen bei Bad Kosen , 8.V.1986, F. Hieke, 1 ex ( ZMHB) ; Kosterbeck near Rostock , 13.VI.1982, M. Schülke, 2 exx ( ZMHB) ; Leipzig , 11.VI.1904, Linke, 1 ex ( MTD) ; 23.VI.1907, 1 ex ( MTD) ; 4.VIII.1907, 8 exx ( MTD) ; 25.VIII.1907, 2 exx ( MTD) ; 12.IX.1909, 1 ex ( MTD) ; 20.V.1917, 1 ex ( MTD) ; 30.VII.1917, 2 exx ( MTD) ; 18.V.1918, 1 ex ( MTD) ; 13.VI.1926, 1 ex ( MTD) ; 19.VIII.1928, 1 ex ( MTD) ; 12.VIII.1934, 1 ex ( MTD) ; 25.VII.1939, 1 ex ( MTD) ; Leipzig , 17.IX.1944, 2 exx ( MTD) ; 6.X.1961, Dorn, 8 exx ( ZMHB) ; Lodderitz, Elbe, Sachs.­Anhalt. , 2.VI.1991, J. Schulze, 4 exx ( ISEA) ; 23 exx ( ZMHB); Luckenwalde , 1 ex ( DEI) ; Mosigkau , 1.VII.1909, 1 ex ( MTD) ; Muggelsee near Rahnsdorf , 1 ex ( MIZ) ; Nahmitz , 1 ex ( DEI) ; Nauen , 29.V.1944, 1 ex ( ZMHB) ; east bank of Muritz, Muritzhof, Muritzufer an der Spuklochkoppel , 22.VI.1991, M. Uhlig, 2 exx ( ZMHB) ; Plaue , Thuringia, 30.VII.1950, W. Liebmann, 1 ex ( DEI) ; Rahnsdorf , 1 ex ( DEI) ; Rheinauen am Kaiserstuhl near Burkheim , 29.VIII.1962, V. Puthz, 15 exx ( MHNG) ; Rostock , 1 ex ( ZMUC) ; Strausberg environs, 1 ex ( DEI) ; Ulm , 1 ex ( NMPC) ; Vissel , 15.V.1927, 1 ex ( MNHN) ; Wannsee, Cladow , 3 exx ( DEI) ; Waren, Miritzhof, NSG "Ostufer der Muritz", Muritz river , 16.VI.1976, M. Schülke, 2 exx ( ZMHB) ; Waren, NSG "Ostufer der Muritz", Muritz river , 23.V.1983, M. Uhlig, 1 ex ( ZMHB) ; 25.VII.1987, M. Uhlig, 1 ex ( ZMHB) ; Worlitz , 1 ex ( CAS) ; 2 exx ( ZMHB); Wurttemberg , 9.VI.1909, 3 exx ( ZMHB) ; 7 exx ( MTD); Wurzen , 30.VII.1917, Uhmann, 1 ex ( ZMHB) ; “ Germania ” (localities not specified), 1 ex ( CAS) ; 1 ex ( ZMUC); “ Saxonia ” (localities not specified), 2 ex ( MTD) ; 1 ex ( ZMUC). Holland: Ben Haag, 1 ex ( NMW) ; Ommen, Valck Lucassen , 12.VI.1929, 1 ex ( FMNH) . Hungary: Békés, Gyula, Fekete­Körös , 90 m, 27.V.1980, Ádám, 1 ex ( HNHM) ; Békés, Sarkad, Remiz­erdö , 90 m, 25.V.1982, Ádám, 1 ex ( HNHM) ; Békés, Kétegyháza , 90 m, 25.VI.1983, Ádám, 1 ex ( HNHM) ; Budapest, Margit Sziget , 23.V.1908, Rambousek, 3 exx ( NMPC) ; Bükki National Park, Miskolc, Garadna­volgy , 250 m, 16.IX.1982, Ádám, 1 ex ( HNHM) ; Kiskunsági National Park, Kunfehértó, Fehér­tó , 29.VIII.1979, Hámori, 4 exx ( HNHM) ; Kiskunsági National Park, Lakitelek, Tóserdö , 25.V.1977, L. Tóth, 1 ex ( HNHM) ; Lillafured, Tógazdaság , 5.X.1958, Vásarhelyi, 1 ex ( HNHM) ; Mehadia , 1 ex ( NMPC) ; Sopron, Moczarski , 5 exx ( NMW) ; Vas m., Csörötnek, Rába­part , 7.X.1980, Ádám, 1 ex ( HNHM) ; Vas m., Meggyeskovácsi , 1.VI.1979, Ádám, 5 exx ( HNHM) ; Vas m., Rátot, Rába­part , 100 m, 10.X.1983, Ádám, 3 exx ( HNHM) . Italy: Lago di Idro , 28.VIII.1992, A. Zanetti, 1 ex ( AZPC) ; Eremo , 23.VI.1948, Fagel, 1 ex ( IRSNB) ; Lombardia, Po­Ticino , 8.X.1961, Rosa, 1 ex ( NMW) ; Pavia , 1919, 3 exx ( MHNG) ; Torino , 1 ex ( BMNH) . Poland: Brzesko Nowe , Dr Klička, 1 ex ( NMPC) ; Drogowle , 29.VIII.1918, Klička, 3 exx ( NMPC) ; “ Glatz ” [Kłodzko], Letzner, 4 exx ( DEI) ; Jarosław koło Krakowa , 1 ex ( FMNH) ; Kazimierza, Klička , 14 exx ( NMPC) ; Kielce , 18.V.1918, Klička, 1 ex ( NMPC) ; Łęczyca , 20.V.1935, 1 ex ( MIZ) ; Nowy S cz distr., Stobiecki, 1 ex ( ISEA) ; Przemyśl, Trella , 1 ex ( ISEA) ; Skolimów , 26.V.1935, Tenenbaum, 4 exx ( FMNH) ; Warszawa­Bielany , 17.VI.1889, 5 exx ( USMB) ; 18.V.1891, 2 exx ( USMB) ; 19.IV.1935, Tenenbaum, 1 ex ( FMNH) ; Warszawa­Saska Kępa , 1.V.1901, 4 exx ( USMB) ; 2.X.1901, 2 ex ( USMB) ; Warszawa­ Ṡwider , 3.VI.1902, 2 ex ( USMB) ; Warszawa­Zegrze , 7.VI.1935, Tenenbaum, 2 exx ( FMNH) ; 24.VI.1934, 2 exx ( MIZ) ; “ Zagdansk ”?, Klička, 1 ex ( NMPC) . Romania: Karpaten, Sinaia Prahova­Tal environs, 900–1600 m, 11–17.VIII.1982, M. Schülke, 3 exx ( ZMHB) . Russia: Krasnoyarsk, Yenisey, Gassner, 1 ex ( NMW) ; “ Sarepta " [Volgograd], V. Bodemeyer, 1 ex ( FMNH) ; ” Simbirsk ” [Ulyanovsk], 1917, Jureèek, 2 exx ( NMPC) ; West Sibiria, Barnaul, Ob River , 1918–1920, Frieb, 13 exx ( NMW) . Slovakia: Nove Mesto , 9.V.1937, A. Matejka, 1 ex ( NMPC) ; Sturovo, Donau river , 3.IX.1983, M. Schülke, 2 exx ( ZMHB) ; O. Kavan , 4 exx ( NMPC) . Sweden: Akarp , 1.IV.1948, Axel Olsson, 1 ex ( NHRS) ; Arlov , 2.V.1959, E. Wiren, 1 ex ( MZLU) ; 12.V.1959, 3 exx ( MZLU) ; Boajokloster , 20.VII.1897, O. Moller, 4 exx ( MZLU) ; Dazsnes , 17.VI.1939, Wiren, 1 ex ( MZLU) ; Deja, E. Wiren, 2 exx ( MZLU) ; Greva , 29.V.1950, E. Wiren, 1 ex ( MZLU) ; Hasslarp , 20.V.1963, F. Olsson, 7 exx ( MZLU) ; Hassleholm , 27.VII.1957, Israelson, 3 exx ( MZLU) ; Ivo , 2.VI.1947, A. Sundholm, 1 ex ( MZLU) ; Karlebo sogn., Niva , 11–31.VII.1962, Israelson, 2 exx ( MZLU) ; Letje, Sandin , 2 exx ( MZLU) ; Lund , 11.V.1934, 45 exx ( MZLU) ; Malmo, Palm , 5 exx ( MZLU) ; Matterods , 1.VII.1957, G. Israelson, 6 exx ( MZLU) ; Ortefta , 22.VII.1946, A. Sundholm, 1 ex ( MZLU) ; 5.VI.1947, 1 ex ( MZLU) ; Ravlunda, Klammersback , 2.VII.1948, Bo Tjeder, 1 ex ( MZLU) ; Revinge , 5.VI.1947, A. Sundholm, 1 ex ( MZLU) ; Varnamo , 3.VI.1936, 3 exx ( MZLU) ; Vinslov , 1.VII.1958, Israelson, 2 exx ( MZLU) ; Vitaby , 3–16.VII.1936, O. Sjoberg, 1 ex ( NHRS) . Switzerland: Aarberg, Kt. Bern, VIII.1937, 1 ex ( ETH) ; Dhbendorf , 4.V.1862, 1 ex ( ETH) ; Faoug , Kt. Waad, VII.1950, 1 ex ( ETH) ; Gampelen, Kt. Bern, VIII.1936, 1 ex ( ETH) ; Hindelbank, Kt. Bern, V.1969, 1 ex ( ETH) . Ukraine: Zaleszczyki , 3.VIII.1930, Tenenbaum, 1 ex ( FMNH) .

Redescription. Body. Length 2.7–3.5 mm, convex, parallel­sided, glossy; body colour pitchy brown to black; legs and antennae reddish brown to brown, three basal tergites often paler, brownish red, head and pronotum with leaden reflection.

Head circular in outline, convex, eyes large, moderately protruding from lateral contours of head, length of each seen from above subequal to that of postocular region; surface of head with isodiametric mesh microsculpture; puncturation fine and moderately dense. Antennae clearly increasing in width apically, antennomere 3 shorter than 2, antennomeres 4–8 longer than wide, antennomeres 9–10 sexually dimorphic, quadrate in male and slightly transverse in female, antennomere 11 nearly conical.

Pronotum quadrate, lateral sides gradually narrowed in straight line to obtuse hind angles; before base with small and shallow transverse impression; surface with fine isodiametric mesh microsculpture; puncturation fine, dense and asperate; pubescence at midline directed posteriorly.

Elytra subquadrate, at suture as long as pronotal length at midline; surface lacking microsculpture; puncturation fine, dense and asperate.

Abdomen parallel­sided; bases of tergites 3–5 each with deep transverse impression, impressions with 7–8 longitudinal ridges and distinct median carina; tergal puncturation fine and moderately dense, punctures moderately small and well visible, surface without microsculpture.

Male. Aedeagus as in Figs. 123–124.

Female. Spermatheca as in Fig. 125.

Taxonomic notes. Tachyusa concinna was synonymised by Fauvel (1895) with T. coarctata . The species here treated as T. concinna is in agreement with the original description and the later redescription of Mulsant & Rey (1875). Heers collection does not exist, therefore, in the interest of stability of nomenclature and according to ICZN (1999: Article 75) a neotype is designated.

Remarks. Tachyusa concinna is closely related to Tachyusa harbinica from which it can be distinguished by the stouter body, the more convex pronotum, the tergite 8 without microsculpture, the tergal basal transverse impressions with median carina and by the shape of the aedeagus.

Distribution. Tachyusa concinna is a Western Palaearctic species distributed from France and Italy in the west to the Yenisey River in the east.