Permula Handlirsch, 1919

Aristov, Daniil S. & Storozhenko, Sergey Yu., 2011, Review of the Permian family Permulidae nomen novum pro Aliculidae Storozhenko, 1997 (Grylloblattida), ZooKeys 130, pp. 111-130 : 113-114

Permula Handlirsch, 1919 nom. resurr.

Alicula Schlechtendal 1913: Pl. 2, Figs 10a, b (type species: Alicula lebachensis Schlechtendal, 1913, by monotypy); Carpenter 1992: 109 (partim); Storozhenko 1998: 97 (partim). Invalid name according to Article 56 of the Code (ICZN, 1999).

Permula Handlirsch 1919: 32 (type species: Permula lebachensis Handlirsch, 1919, by original designation); Handlirsch 1922: 77; Kukalová 1964: 46, 47 (partim).

Description.

The base of costal area in forewing broadly rounded; costal area is considerably broader than subcostal one, crossed with simple or furcated anterior branches of SC; RA simple; RS with more than two branches; the base of M distinctly separated from CuA; M forked before the base of RS; CuA with 2-4 branches, first fork of CuA is situated proximal to fork of M; the base of anterior branch of CuA sclerotized; CuP curved or straight; A1 and A2 simple.

Composition.

Four species from the Lower and Middle Permian of Europe.

Notes.

Apparently being unaware of Schlechtendal’s paper, Handlirsch (1919) described Permula lebachensis as a new genus and species based on the same specimen as Alicula lebachensis Schlechtendal, 1913. Later Carpenter (1992) synonymized Permula lebachensis Handlirsch, 1919 with Alicula lebachensis Schlechtendal, 1913 and included Permula acra Kukalová, 1964 in the genus Alicula . But the generic name Alicula Schlechtendal, 1913 is a junior homonym of Alicula Eichwald, 1830 and must be replaced by Permula Handlirsch 1919. Herein two species are transferred from Sojanopermula to Permula based on the shape of costal area and one new species is described. Moreover, Permula acra Kukalová, 1964 is not congeneric with other species of Permula and is placed in a new genus of the family Ideliidae (see below).

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