Cecidomyia dubia

Kolesik, Peter & Gagné, Raymond J., 2016, Revision of early taxa of Australian gall midges (Diptera: Cecidomyiidae), Zootaxa 4205 (4), pp. 301-338 : 330-336

Nomina dubia

The condition of the type specimens is not given for the species of Micromyinae in the list that follows. Those specimens were studied by Jaschhof (2010) and returned separately to ANIC.

adusta Skuse, 1888: 82 , Cecidomyia (Diplosis) . Holotype male, “Woronora (Masters and Skuse). October”, in ANIC. The pinned type lacks its head and abdomen. Skuse (1890: 378) corrected himself and stated that his original type was a female and not a male, but that is not possible to check now. He then described the male on the basis of additional dried specimens still in the Skuse pinned collection in ANIC but we cannot determine now that they are the same species as his original specimen. Placed with nomina dubia by Gagné (2004: 283).

albulipennis Skuse, 1890: 378 , Cecidomyia (Diplosis) . Syntypes, male(s) and female(s), “ Sydney (Skuse); Dunoon , Richmond River , NSW (Helms); Burpengary, near Brisbane, Queensland (Dr. T. L. Bancroft), one specimen in Coll . Queensland Museum. February to April.” In ANIC, we found three specimens: one labelled “ Dunoon " and two without locality data, none of the three with its abdomen. The specimen collected at Burpengary is not part of the Queensland Museum collection, and is presumably lost. Placed with nomina dubia by Gagné (2004: 283).

amplipennis Skuse, 1888: 142 , Campylomyza . Declared a nomen dubium by Jaschhof (2010: 37) . No insects were attached to the pins in NHMW.

araneosa Skuse, 1888: 82 , Cecidomyia (Diplosis) . Holotype female, “ Elizabeth Bay (Skuse). October “, in ANIC . In the original description the type is stated as male but Skuse (1890: 378) corrected himself and wrote that his original type was a female. The mounted holotype retains its wings, shrivelled postabdomen that appears barely protrusible, with two large cerci, one 4-segmented palpus, one antenna with 12 flagellomeres, but no legs. It belongs to the Cecidomyiidi but lacks distinctive morphological characters on preserved body parts to place it further. This species was moved to “ Unplaced Cecidomyiinae ” in Gagné (2004). We now consider it a nomen dubium. New status.

ardens Skuse, 1888: 75 , Cecidomyia (Diplosis) . Holotype female, “Sydney (Skuse). January“, in ANIC. The pinned specimen lacks its head and abdomen. Placed with nomina dubia by Gagné (2004: 283).

aurata Skuse, 1888: 129 , Lasioptera . Holotype male, “Middle Harbour (Skuse). September”, in ANIC. The mounted specimen retains its terminalia and a claw and fits Lasioptera , but does not show special characters to distinguish it from other congeners. We consider this species a nomen dubium. New status.

bellula Skuse, 1888: 87 , Cecidomyia (Diplosis) . Holotype male, “Peat’s Ferry (Masters and Skuse)“, in ANIC. The pinned type lacks its abdomen. Placed with nomina dubia by Gagné (2004: 283).

bombycina Skuse, 1888: 79 , Cecidomyia (Diplosis) . Holotype female, “Sydney (Skuse)“, in ANIC. Only parts of wings and legs remain of the pinned type. Placed with nomina dubia by Gagné (2004: 283).

brevipennis Skuse, 1888: 71 , Cecidomyia (Diplosis) . Holotype male, “Elizabeth Bay (Masters). December“, in ANIC. The mounted type retains its wings, antennae with 12 binodal flagellomeres bearing one circumfilum on proximal and two on distal node, partially shrivelled head with occipital protuberance, 4-segmented palpi, simple tarsal claws, empodia as long as claws, pulvilli half length of claws, shrivelled terminalia with simple gonocoxites bearing no mediobasal lobes, distally tapered aedeagus and basally microtrichose gonostyli. It belongs to Cecidomyiidi but it has damaged terminalia and lacks distinctive morphological characters on other body parts to compare it to males from other genera and species. Moved to "Unplaced Cecidomyiinae " in Gagné (2004: 275), we now consider this species a nomen dubium. New status.

brevitata Gagné, 1989: 161 , Cecidomyia , new name for Cecidomyia (Diplosis) abbreviata Skuse 1888: 75 by Gagné (1989b). Preoccupied by Cecidomyia abbreviata Waltl 1837: 279 . Holotype female, “Sydney (Skuse). November“, in ANIC. The pinned type lacks the abdomen. Placed with nomina dubia by Gagné (2004: 283).

caeca Skuse, 1888: 76 , Cecidomyia (Diplosis) . Holotype male, “ Elizabeth Bay (Skuse)“, in ANIC . Only legs of the pinned type remain. Skuse (1890: 377) identified five further specimens as this species: “Five old specimens recently found in the collection of the late Mr. W. S. Macleay belong to this species [ Cecidomyia (Diplosis) caeca ], and are labelled “ Cecidomyia muscorum ”; on dead insects, N.S.W. I know nothing of this habit from personal observation.”. We have not found these specimens in ANIC. “ Cecidomyia muscorum ” was placed with nomina nuda by Gagné (2004: 286). Cecidomyia caeca was added to nomina dubia by Gagné (2004: 283).

certa Skuse, 1888: 100 , Cecidomyia (Diplosis) . Holotype female, “Woronora (Masters and Skuse)”, in ANIC. The abdomen of the pinned type is partly destroyed. Placed with nomina dubia by Gagné (2004: 283).

cineraria Skuse, 1888: 106 , Cecidomyia (Diplosis) . Holotype male, “Woronora, Sydney (Masters and Skuse). October”, in ANIC. The pinned type lacks its antennae and abdomen. Placed with nomina dubia by Gagné (2004: 283).

conferta Skuse, 1888: 118 , Cecidomyia (Epidosis) . Holotype female, “ Elizabeth Bay (Masters)”, in ANIC. Only part of a leg remains of the type. Placed with nomina dubia by Gagné (2004: 283).

confinis Skuse, 1888: 70 , Cecidomyia (Diplosis) . Holotype male, “Sydney (Skuse). January“, in ANIC. The pinned type lacks its antennae and abdomen. Placed with nomina dubia by Gagné (2004: 283).

conspecta Skuse, 1888: 91 , Cecidomyia (Diplosis) . Holotype female, “ Richmond , Hawkesbury district (Skuse). January ”, in ANIC. The pinned type lacks its abdomen. Placed with nomina dubia by Gagné (2004: 283).

contigua Skuse, 1888: 103 , Cecidomyia (Diplosis) . Holotype female, “ Wheeny Creek , Hawkesbury district (Skuse). January ”, in ANIC. No specimen remains attached to the pinned card. Placed with nomina dubia by Gagné (2004: 283).

corusca Skuse, 1888: 131 , Lasioptera . Syntype males, “Gosford (Skuse). February”, in ANIC. Of six males three are substantially damaged and left pinned, three were mounted. Mounted specimens have most characters preserved but lack special characters to distinguish them from other Lasioptera males. We place this species here with nomina dubia . New status.

crocea Skuse, 1888: 136 , Campylomyza . Declared a nomen dubium by Jaschhof (2010: 37) .

dibapha Skuse, 1888: 98 , Cecidomyia (Diplosis) . Syntype females, “ Illawarra district (Masters and Skuse); Coogee Bay (Skuse). September ”, in ANIC . Of the two pinned specimens in ANIC, we did not mount the one from Coogee Bay as it lacks its head and abdomen and has a wing colour pattern different from the other. The syntype from Illawarra district , which we mounted, retains all body parts except the tarsi. Assigned to unplaced Cecidomyiinae by Gagné (2004: 276), it belongs to Contarinia but lacks distinctive morphological characters to distinguish it from other congeneric females. We declare it a nomen dubium. New status.

difficilis Skuse, 1888: 63 , Cecidomyia . Holotype female, “Elizabeth Bay (Masters). Beginning of January”, in ANIC. The mounted type specimen has preserved wings, damaged head capsule, 4-segmented palpi, antennae with 11 flagellomeres, the terminal twice as long as others, toothed tarsal claws, empodia as long as claws, preserved terminalia with eighth tergite split into two longitudinal sclerites, remaining body parts missing. This species belongs to Dasineura but lacks distinctive morphological traits to separate it from congeneric females. We move it here to nomina dubia . New status.

discedens Skuse, 1888: 93 , Cecidomyia (Diplosis) . Holotype female, “ Elizabeth Bay (Masters)”, in ANIC. The pinned type lacks its head and abdomen. Placed with nomina dubia by Gagné (2004: 283).

distenta Skuse, 1888: 115 , Cecidomyia (Epidosis) . Holotype female, “ Elizabeth Bay (Skuse). Beginning of December ”, in ANIC. The type has shrivelled antennae and collapsed terminalia. It belongs to Porricondylinae but cannot be compared unambigiously to other species. Assigned to unplaced Porricondylinae by Gagné (2004: 63). We move it here to nomina dubia . New status.

erronea Skuse, 1888: 94 , Cecidomyia (Diplosis) . Holotype female, “Sydney (Skuse). End of November”, in ANIC. The pinned type lacks its abdomen and the collection data label. Assigned to unplaced Cecidomyiinae by Gagné (2004: 277). Here we declare it a nomen dubium as the type cannot be properly compared to females of other species. New status.

exigua Skuse, 1888: 122 , Cecidomyia (Epidosis) . Holotype male [considered a female by Skuse], “ Glenbrook , Blue Mountains (Masters)”, in ANIC. The type has shrivelled terminalia. It belongs to Porricondylinae but cannot be compared unambigiously to other species. Assigned to unplaced Porricondylinae by Gagné (2004: 63). We move it here to nomina dubia . New status.

faceta Skuse, 1888: 81 , Cecidomyia (Diplosis) . Holotype male, “Sydney (Skuse)“, in ANIC. The type is missing from its pinned card. Placed with nomina dubia by Gagné (2004: 283).

fallax Skuse, 1888: 85 , Cecidomyia (Diplosis) . Holotype female, “Woronora Masters and Skuse)”, not in ANIC, presumed lost. Skuse (1890: 382) declared Cecidomyia (Diplosis) fallax a junior synonym of C. negotiosa , which itself can no longer be identified. Placed with nomina dubia by Gagné (2004: 284).

flammula Skuse, 1888: 124 , Cecidomyia (Asynapta) . Holotype female, “Sydney (Masters)”, in ANIC. The end of the abdomen of the pinned type is destroyed. Placed with nomina dubia by Gagné (2004: 283).

frequens Skuse, 1888: 99 , Cecidomyia (Diplosis) . Syntypes, male and female, “Elizabeth Bay (Masters and Skuse). End of December”, in ANIC. Both types were mounted. The male has wings, head with 4-segmented palpi and occipital protuberance but without flagellomeres, legs with simple claws as long as empodia, and distorted terminalia with aedeagus missing. The female has all parts preserved, but with the ovipositor inside the abdomen and not properly examinable. This species was assigned to unplaced Cecidomyiinae by Gagné (2004: 277). It belongs to Contarinia but the types lack distinctive morphological characters on preserved body parts that can distinguish them from congeners. We place it with nomina dubia . New status.

furva Skuse, 1888: 90 , Cecidomyia (Diplosis) . Holotype female, “Middle Harbour (Skuse)”, in ANIC. The pinned type lacks its abdomen. Placed with nomina dubia by Gagné (2004: 284).

gibberosa Skuse, 1888: 123 , Cecidomyia (Epidosis) . Holotype female, “ Elizabeth Bay (Masters)”, in ANIC. Although Skuse described a “specimen” the pinned type card bears two specimens but neither has an abdomen. Placed with nomina dubia by Gagné (2004: 284).

gibbula Skuse, 1888: 67 , Cecidomyia . Holotype female, “Middle Harbour (Skuse) “, in ANIC. The pinned type lacks its abdomen. Placed with nomina dubia by Gagné (2004: 284).

gilva Skuse, 1888: 93 , Cecidomyia (Diplosis) . Holotype male, “Gosford (Skuse)”, in ANIC. The pinned type lacks the abdomen and antennae. Placed with nomina dubia by Gagné (2004: 284).

Gonioclema Skuse, 1888: 60 , as subgenus of Cecidomyi a. Type species, Cecidomyia pauxillula Skuse, 1888 (monotypy). Skuse (1888: Fig. 3 ) illustrated the wing showing R4+5 straight and joining C at wing apex, Rs closer to distal end of R1 than arculus, Cu simple and fading apically, wingfold fading basally and apically. He described female flagellomeres as subcylindrical, 11 in number. The description is ambiguous and the holotype has been mostly destroyed by insects (see Cecidomyia pauxillula listing in nomina dubia ). Placed with nomina dubia by Gagné (2004: 282).

gracilis Skuse, 1888: 117 , Cecidomyia (Epidosis) : see listing of Porricondyla skusei Kieffer.

grandipennis Skuse, 1890: 384 , Cecidomyia (Epidosis) . Holotype male, “ Mossman’s Bay , near Sydney (Skuse). One specimen in September ”, in ANIC. The pinned type lacks its antennae and abdomen. Placed with nomina dubia by Gagné (2004: 284).

helvipes Skuse, 1888: 132 , Lasioptera . Holotype female [mistaken for male by Skuse], “Sydney (Masters)”, in ANIC. Female mounted: all body parts present except for legs, but ovipositor is shrivelled inside collapsed remains of abdomen; hooked setae, typical for Lasioptera , are present on cerci but their number and position are obscured. No other characters will allow one to distinguish the type from congeneric females. We move it here to nomina dubia . New status.

helmsi Skuse, 1890: 379 , Cecidomyia (Diplosis) . Syntypes, males and females, “ Dunoon , Richmond River , NSW (Helms). Twenty-two specimens in March and April ”, in ANIC. The pinned types are all largely destroyed and none bears an abdomen. Placed with nomina dubia by Gagné (2004: 284).

humilis Skuse, 1888: 95 , Cecidomyia (Diplosis) . Syntypes, male and female, “ Wheeny Creek , Hawkesbury district (Skuse). January ”, in ANIC. Of the three pinned specimens glued to one card none retains its head or abdomen. Placed with nomina dubia by Gagné (2004: 284).

indotata Skuse, 1888: 107 , Cecidomyia (Diplosis) . Holotype male, “Middle Harbour (Skuse). In November”, in ANIC. The pinned type lacks its abdomen. Placed with nomina dubia by Gagné (2004: 284).

loewi Skuse, 1888: 108 , Cecidomyia (Asphondylia) . Holotype male [mistaken for female by Skuse (1888)], “Como (Masters and Skuse). September”, in ANIC. The pinned type is a male, a probable Asphondylia , whose abdomen could have appeared to be that of a female to Skuse but it now lacks its head and can never be identified with certainty. Skuse (1890: 382) later described a male which he associated with this species, but that specimen is presumably lost. Placed with nomina dubia by Gagné (2004: 284).

lucida Skuse, 1888: 73 , Cecidomyia (Diplosis) . Holotype male, “Woronora (Masters and Skuse)”, in ANIC. The pinned specimen lacks its abdomen. Placed with nomina dubia by Gagné (2004: 284).

lutulenta Skuse, 1888: 110 , Cecidomyia (Hormomyia) . Holotype male, “ Elizabeth Bay (Skuse)”, in ANIC. The pinned type lacks its abdomen. Placed with nomina dubia by Gagné (2004: 284).

macella Skuse, 1888: 121 , Cecidomyia (Epidosis) . Holotype female, “Sydney (Skuse)”, in ANIC. Nothing remains of the pinned type on the card. Placed with nomina dubia by Gagné (2004: 284).

macleayi Skuse, 1888: 97 , Cecidomyia (Diplosis) . Holotype female, “Woronora (Masters and Skuse). October; in caves”, in ANIC. The mounted type retains a wing, head with 4-segmented palpi, occipital protuberance, antenna with 12 barrel-shaped flagellomeres, slightly narrowed at midlength, circumfila as sparse network of low loops, legs with claws (large, toothed, bent at basal third, empodia reaching bend in claws) and terminalia with large cerci bearing long simple setae and wide hyporoct with pair of short setae. This species was assigned to unplaced Cecidomyiinae by Gagné (2004: 279). It belongs to Cecidomyiidi but the type lacks special morphological characters that would distinguish it from other species. We move it here to nomina dubia . New status.

magnifica Skuse, 1888: 116 , Cecidomyia (Epidosis) . Holotype male, “ Elizabeth Bay (Masters). January ”, in ANIC. The type has shrivelled terminalia. It belongs to Porricondylinae but cannot be compared unambigiously to other species. Assigned to unplaced Porricondylinae by Gagné (2004: 63). We move it here to nomina dubia . New status.

mastersi Skuse, 1888: 127 , Lasioptera . Holotype female, “Woronora (Masters and Skuse). October”, in ANIC. The head and postabdomen of the pinned type are missing. Placed with nomina dubia by Gagné (2004: 284).

mastersi Skuse, 1888: 59 , Miastor . Holotype male, “ Elizabeth Bay (Skuse). February ”, in ANIC. Nothing remains of the type on its card. Placed with nomina dubia by Gagné (2004: 64).

mollipes Skuse, 1888: 92 , Cecidomyia (Diplosis) . Syntypes, male and female, “Sydney (Masters and Skuse)”, in ANIC. Neither type bears a locality label. We mounted both types. The male retains a wing, head with occipital protuberance, one palpus (with 4 segments), both antennae with scape and pedicel and first flagellomere on one (consisting of two nodes, one looped circumfilum on proximal and two on distal node), legs with claws (simple, thin, bent beyond midlength, longer than empodia), but is misssing the terminalia. The female has retained its wings, head with occipital protuberance, 4-segmented palpus, antennae with 12 flagellomeres, terminalia with large cercus (one missing). This species was assigned to unplaced Cecidomyiinae by Gagné (2004: 279). It belongs to Cecidomyiidi but the male type lacks terminalia and there are no special morphological characters on surviving body parts of either type that would distinguish it from other species. We place it here with nomina dubia . New status.

montana Skuse, 1888: 84 , Cecidomyia (Diplosis) . Holotype female, “Glenbrook (Masters). January“, in ANIC. Only legs of the pinned type remain. Placed with nomina dubia by Gagné (2004: 284).

Necrophlebia Skuse, 1888: 111 , as subgenus of Cecidomyi a. Type species, Cecidomyia volitans Skuse, 1888 (monotypy). Skuse (1888: Fig. 10 ) illustrated the wing showing R4+5 apically curved and joining C posteriad of the wing apex, Rs halfway between distal end of R1 and arculus, Cu simple and reaching wing margin, wingfold not visible. He described female flagellomeres as subcylindrical, 12 in number. This subgenus cannot be identified because the original description lacks important diagnostic characters and the pinned types of the only included species are damaged beyond recognition (see Cecidomyia volitans listing in nomina dubia ). Placed with nomina dubia by Gagné (2004: 282).

negotiosa Skuse, 1888: 104 , Cecidomyia (Diplosis) . Syntypes, male and female, “ North Willoughby (Masters and Skuse). Beginning of December ”, in ANIC. Parts of eight pinned specimens without abdomens remain on four separate cards. Cecidomyia negotiosa placed with nomina dubia by Gagné (2004: 284).

nobilis Skuse, 1888: 66 , Cecidomyia . Holotype male, “Gosford (Skuse). February”, in ANIC. The pinned type lacks its abdomen. Placed with nomina dubia by Gagné (2004: 284).

nubilipennis Skuse, 1890: 375 , Cecidomyia . Holotype female [mistaken for male by Skuse], “ Elizabeth Bay , near Sydney (Skuse). October. A single specimen taken on a window”, in ANIC . The type lacks the head but retains the thorax with three legs, and the abdomen with terminalia. In Notes and Exhibits of the Linnean Society of NSW (Anonymous 1893) this species is commented on: “Mr. Froggatt exhibited specimens of the galls of Cecidomyia nubilipennis, Sk. , previously unrecorded, from Flemington [NSW], on the leaf-stalkes of Eucalyptus sideriphloia , and of the gnats bred therefrom”. Froggatt’s specimens were not found in ANIC. In view of the level of diagnostic capability at the time of Froggatt’s exhibition, we doubt that his specimens had more than a superficial resemblance to the holotype of C. nubilipennis that was taken on a window. This species was assigned to unplaced Cecidomyiinae by Gagné (2004: 280). We note here that it belongs to Asphondylia . Given that Asphondylia females mostly lack diagnostic characters to allow comparison with congeners, we place C. nubilipennis with nomina dubia . New status.

obsoleta Skuse, 1888: 83 , Cecidomyia (Diplosis) . Holotype female, “Glenbrook (Masters). January”, in ANIC. Only legs of the pinned type remain. Placed with nomina dubia by Gagné (2004: 284).

opipara Skuse, 1888: 118 , Cecidomyia (Epidosis) . Holotype female, “ Elizabeth Bay (Masters). January ”, in ANIC. The type lacks the ovipositor. It belongs to Porricondylinae but cannot be compared unambigiously to other species. Assigned to unplaced Porricondylinae by Gagné (2004: 63). We move it here to nomina dubia . New status.

parietina Skuse, 1888: 125 , Cecidomyia (Asynapta) . Holotype female, “ Elizabeth Bay (Masters and Skuse). January ”, in ANIC. The abdomen of the pinned type is severely damaged. Placed with nomina dubia by Gagné (2004: 285).

parilis Skuse, 1888: 87 , Cecidomyia (Diplosis) . Holotype female, “Middle Harbour (Skuse)”, in ANIC. State of the holotype: wings well preserved; head shrivelled, one antenna with several flagellomeres present, palpi missing; only one mid- and one hindleg present; abdomen present with terminalia folded inside and details obscured. Skuse (1890: 380, Pl. XVI., Fig. 3 [ Fig. 2 h]) assigned to this species four females (mounted by us) that were “Bred from irregular-oval, reddish-brown blisters, 3 to 4 mm. long, having an ochreous-yellow or pale brownish slit (1 to 1½ mm. in length) in the middle; occurring very numerously on the upper side of the leaves of Eucalyptus corymbosa , obtained by Mr. Froggatt, in November, at Weverly, near Sydney. The blisters extend through the leaf, being represented on the under side by very slightly raised brown patches. As many as forty or fifty blisters sometimes occur in one leaf; they are usually dispersed over the surface, but occasionally form clusters of three or four. It appears that the larvae abandon the blisters before assuming their next state; the presence of the longitudinal opening also supports this conclusion.” The four females taken from galls lack the terminalia but have well preserved heads with flagellomeres and wings, and one specimen has claws on at least one of each pair of legs. Their Rs vein lies midway between the arculus and the end of R1 and their claws are all toothed, unlike the holotype of C. parilis that has the Rs vein closer to the arculus and untoothed claws. Because of these differences it is clear that Skuse’s 1890 gall association was incorrect and the gallmaker of Eucalyptus corymbosa remains unnamed. Cecidomyia parilis is assigned here to nomina dubia . New status.

paula Skuse, 1888: 74 , Cecidomyia (Diplosis) . Holotype female, “ Elizabeth Bay (Skuse). End of January ”, in ANIC. Nothing remains of the pinned specimen on the card. Placed with nomina dubia by Gagné (2004: 285).

pauxillula Skuse, 1888: 61 , Cecidomyia (Gonioclema) . Holotype female, “ Elizabeth Bay (Skuse). September ”, in ANIC. Only a wing and a fragment of the thorax remain of the pinned type. Placed with nomina dubia by Gagné (2004: 285).

percita Skuse, 1888: 72 , Cecidomyia (Diplosis) . Holotype female, “Elizabeth Bay (Skuse). December“, in ANIC. The mounted type retains the wings, one foreleg with toothed claws and empodium that is longer than the claws, shrivelled terminalia with evidently barely protrusible large cerci, head with 12 flagellomeres, occipital protuberance and 4-segmented palpi. It belongs to Cecidomyiidi but lacks distinctive morphological characters to compare it to females from other genera and species. We move it here to nomina dubia . New status.

perexilis Skuse, 1888: 119 , Cecidomyia (Epidosis) . Holotype male, “Elizabeth Bay (Skuse)”, in ANIC. The terminalia of the type are too poorly displayed for examination. It belongs to Porricondylinae but cannot be compared unambigiously to other species. Assigned to unplaced Porricondylinae by Gagné (2004: 63), it is moved here to nomina dubia . New status.

perpallida Skuse, 1888: 134 , Campylomyza . Declared a nomen dubium by Jaschhof (2010: 37) .

persimilis Skuse, 1888: 136 , Campylomyza . Declared a nomen dubium by Jaschhof (2010: 37) .

prisca Skuse, 1888: 125 , Cecidomyia (Asynapta) . Holotype female, “ Elizabeth Bay (Masters). Beginning of December ”, in ANIC. Only parts of a wing and a leg remain of the type. Placed with nomina dubia by Gagné (2004: 285).

probata Skuse, 1888: 77 , Cecidomyia (Diplosis) . Holotype female, “Middle Harbour, near Sydney (Masters and Skuse). August”, in ANIC. Only legs of the pinned type remain. Placed with nomina dubia by Gagné (2004: 285).

procax Skuse, 1888: 58 , Miastor . Holotype female, “ Elizabeth Bay , near Sydney (Skuse). Beginning of January ”, in ANIC. Nothing remains of the type on its card. Placed with nomina dubia by Gagné (2004: 64).

quaesita Skuse, 1888: 90 , Cecidomyia (Diplosis) . Holotype female, “Woronora (Masters and Skuse)”, in ANIC. Although the original description was based on a “dried specimen”, two pinned specimens after this name bear the same collection label. Both specimens lack abdomens. Placed with nomina dubia by Gagné (2004: 285).

regilla Skuse, 1888: 64 , Cecidomyia . Holotype female, “ Elizabeth Bay (Skuse). End of January ”, in ANIC. The pinned type has wings similar to those of a Brachineura but lacks the head and abdomen. It can never be identified with certainty. Placed with nomina dubia by Gagné (2004: 285).

rubicunda Skuse, 1888: 109 , Cecidomyia (Asphondylia) . Holotype male, “ Elizabeth Bay (Masters). December ”, in ANIC. The pinned type lacks its abdomen. Placed with nomina dubia by Gagné (2004: 285).

rusticula Skuse, 1888: 89 , Cecidomyia (Diplosis) . Holotype male, “Como (Skuse)”, in ANIC. The pinned type lacks its abdomen. Placed with nomina dubia by Gagné (2004: 285).

saxatilis Skuse, 1888: 102 , Cecidomyia (Diplosis) . Syntypes, male and female, “ Elizabeth Bay , Sydney (Masters and Skuse)”, in ANIC. Neither pinned specimen has a collection data label and only non-diagnostic fragments remain of both. Placed with nomina dubia by Gagné (2004: 285).

scelesta Skuse, 1888: 96 , Cecidomyia (Diplosis) . Holotype male, “ Elizabeth Bay (Skuse). January ”, in ANIC. The pinned type lacks its antennae and abdomen. Placed with nomina dubia by Gagné (2004: 285).

scenica Skuse, 1888: 72 , Cecidomyia (Diplosis) . Holotype male, “ Elizabeth Bay (Masters). September “, in ANIC. The pinned specimen lacks its abdomen. Placed with nomina dubia by Gagné (2004: 285).

senilis Skuse, 1888: 88 , Cecidomyia (Diplosis) . Holotype female, “ Coogee Bay , near Sydney (Skuse). In September ”, in ANIC. Only a wing and leg fragments remain of the pinned type. Placed with nomina dubia by Gagné (2004: 285).

sidneyensis Schiner, 1868: 9 , Campylomyza . Declared nomen dubium by Jaschhof (2010: 36) . No insects were attached to the pins in NHMW.

skusei Kieffer 1913: 273 , Porricondyla . New name for Cecidomyia gracilis Skuse by Kieffer

(1913 b).

Cecidomyia (Epidosis) gracilis Skuse, 1888: 117 . Holotype male, “ Elizabeth Bay (Skuse)”, lost. Assigned to unplaced Porricondylinae by Gagné (2004: 63). We move it here to nomina dubia . New status. Porricondyla exilis Gagné 1989b: 156 , unnecessary new name for Cecidomyia gracilis (Skuse) .

sulfurea Skuse, 1888: 86 , Cecidomyia (Diplosis) . Holotype female, “Sydney (Masters and Skuse)”, in ANIC. The pinned type has no collection label and two legs are all that remains. Placed with nomina dubia by Gagné (2004: 285).

sydneyensis Skuse, 1890: 389 , Lestremia . Holotype female, “Sydney (J. D. Ogilby). One specimen in June”. Apart from that the type is presumably lost, Lestremia females cannot be unambiguously identified to species (M. Jaschhof, pers. comm., 2014).

vegrandis Skuse, 1888: 78 , Cecidomyia (Diplosis) . Syntypes, male and female. “ Elizabeth Bay (Masters and Skuse). January ”, in ANIC. Of the two pinned cards, nothing remains on one and only non-diagnostic fragments are left on the other. Placed with nomina dubia by Gagné (2004: 285).

vicina Skuse, 1888: 138 , Campylomyza . Declared a nomen dubium by Jaschhof (2010: 38) .

villosa Skuse, 1888: 80 , Cecidomyia (Diplosis) . Holotype male, “ Elizabeth Bay (Masters). January ”, in ANIC. Only a wing and leg fragments remain of the pinned type. Placed with nomina dubia by Gagné (2004: 285).

virgata Skuse, 1890: 386 , Lasioptera . Syntypes, male and female, “ Elizabeth Bay (Skuse). November ”, in ANIC. Both pinned types are substantially damaged and lack their abdomens. Placed with nomina dubia by Gagné (2004: 285).

volitans Skuse, 1888: 111 , Cecidomyia (Necrophlebia) . Syntype females, “Middle Harbour (Skuse) and Woronora (Masters)”, in ANIC. The head and wings are missing from the pinned Middle Harbour specimen and only museum pest frass remains on the card of the Woronora specimen. Placed with nomina dubia by Gagné (2004: 285).

wildi Skuse, 1890: 387 , Lasioptera . Syntypes, male and female, “Botanical Gardens, Brisbane, Q. (C. J. Wild). March. Specimens in Coll. Queensland Museum”, in ANIC, although originally indicated as in collection of the Queensland Museum. Only legs remain of the pinned female. The mounted male is well-preserved but shows no special characters to distinguish it from other Lasioptera males. We move it here to nomina dubia . New status.

Figures

FIGURE 2. (a) Galls of Lasioptera miscella Skuse on leaf stalks of Eucalyptus haemastoma, (b) leaf galls of Okriomyia omalanthi (Skuse) on Omalanthus populifolius, (c) stem galls of Daphnephila glandifex Kieffer on Machilus gamblei (Lauraceae) from India which Edwards (1916) stated resembled galls of Asphondylia hilli Edwards, (d) flowers galls of Dasineura dielsi Rübsaamen on Acacia cyclops, (e) galls of Cecidomyia eucalypti Skuse (unplaced Cecidomyiinae) on stems of Eucalyptus haemastoma, (f) galls of Mesodiplosis callitridis Kolesik on branch shoot tips of Callitris endlicheri from which the inquiline Gladiodiplosis frenelae (Skuse) was reared, (g) flower galls of Dasineura acaciaelongifoliae (Skuse) on Acacia longifolia, (h) leaf galls attributed to Cecidomyia parilis Skuse (nomen dubium) on Eucalyptus corymbosa, (i) leaf galls of Heteropeza transmarina Schiner (unplaced Cecidomyiinae) on Callistemon sp., (j) flower and vegetative bud galls of Dasineura frauenfeldi (Schiner) on Leptospermum laevigatum. Left: original figure from Schiner (1868); centre: galls collected with type series in 1858; right: galls collected by Kolesik at Queenscliff, Victoria, in 2008. Figures a, b, e – h from Skuse (1890); c from Kieffer (1913 b); d from Rübsaamen (1916); i, j (left) from Schiner (1868).

FIGURE 3. Asphondylia hilli Edwards. a – e: male, f – g: female, h: pupa. (a) wing, (b) terminalia in dorsal view, (c) last tarsomere with claw and empodium, (d) first tarsomere, (e) palpus, (f) postabdomen in dorsal view, (g) last flagellomeres, (h) antennal horns. Figures g and h from Edwards (1916).

FIGURE 10. Haplusia bella (Skuse). a – d: holotype female, e – f: male caught by Colless in 1981. (a) wing (partially damaged), (b) tarsal claw with empodium, (c) sixth flagellomere, (d) ovipositor in lateral view, (e) sixth flagellomere (setae cut short diagrammatically), (f) terminalia in dorsal view (aedeagus broken off).