Macrohydnobius contortus ( Hatch, 1957 ) Peck & Cook, 2009

Peck, Stewart B. & Cook, Joyce, 2009, Review of the Sogdini of North and Central America (Coleoptera: Leiodidae: Leiodinae) with descriptions of fourteen new species and three new genera, Zootaxa 2102 (1), pp. 1-74 : 44-47

Macrohydnobius contortus ( Hatch, 1957) , new combination

( Figs. 123–130 , 131 )

Hydnobius contortus Hatch, 1957: 24 . Type material. Hydnobius contortus , holotype, male, in SMDV, bearing white label “Creston, B.C. / 12-XII 1951 / G. Stace Smith ”; white handwritten label “on snow”; red handwritten label “TYPE [male symbol] / Hydnobius / contortus / 1954— M. Hatch ”; white label “ Ex. G. Stace-Smith / collection / purchased 1960”; seen and dissected. Type locality: Creston, BC.

Additional material examined. We examined 234 specimens (See Appendix). Diagnosis. Body reddish brown, shining. Length of pronotum + elytra = 4.3–6.4 mm (males), 4.2–6.3 mm (females). Head finely punctate with no microsculpture; with a small median depression and a pair of small lateral depressions anterior to median depression. Pronotum narrow, widest at basal one-third, length:width ratio = 1:1.4; sides obtusely angulate at basal one-third, apical two-thirds weakly concave; basal angles obtuse; with a pair of small basal impressions joined by a transverse, closely punctate, impressed line; finely punctate with microsculpture of broken fine lines. Elytra elongate, wider than pronotum, ratio length:width = 1:0.6; with 9 closely punctate striae; stria 1 impressed, striae 6 and 7 do not reach apex; intervals with 2–3 rows of fine punctures and transversely striolate; uneven intervals with scattered larger punctures. Antennal club ( Fig. 123 ) ratio length:width 1:3.1; width ratio of antennomeres 7:8:9 = 1.4:1:1.5. Mandibles ( Figs. 124, 125 ) elongate, sexually dimorphic. Left mandible in both sexes ( Figs. 124, 125 ) with tooth on outer margin, acute tooth on apical one-half of inner margin. Right mandible of male ( Fig. 124 ) strongly curved apically and lacking tooth on outer margin; in female ( Fig. 125 ) with tooth on outer margin; both sexes with two teeth at middle of inner margin. Male metatrochanter with small blunt tooth before apex. All femora unarmed in both sexes; male protibia ( Fig. 126 ) broad at apex, with large sinuate spur at apex; outer margin spinose; female protibia more slender, outer margin spinose, spur unmodified; mesotibia and metatibia broad apically in male, slender in female, outer margin spinose in both sexes. Male. Aedeagus ( Fig. 127 ) with median lobe elongate, apical one-half strongly narrowed, apex tapering to narrow triangle; parameres broad, wider than adjacent median lobe, concave on apical one-third or more of inner margin, with setae apically. Internal sac ( Fig. 128 ) with scale-like setae; at middle with two distinct areas of dense scale-like setae. Female. Coxites ( Fig. 130 ) broad, flat, with narrow apices curved dorsad; small styli inserted on ventral face of coxites; coxites with setae in area of insertion of styli. Sternite 8 ( Fig. 129 ) deeply emarginate apically; anterior apophysis narrow, parallel-sided.

Distribution. The species occurs in forested northwestern North America eastwards to Utah ( Fig. 131 ). We have seen specimens from Canada: the province of British Columbia; USA: the states of California, Montana, Oregon, and Utah.

Field notes and habitats. Almost all adults have been collected on snow, in presumably forested habitats, and with a small number in rotten wood, in flight, at a light trap, and in a pitfall or flight intercept trap. A single specimen from Cedar Breaks National Monument, Iron Co., Utah was collected from the Fuzzy Truffle , Geopora cooperi , cited by Fogel and Peck (1975) .

Seasonality. Adults have been collected in all months except April and June, with most in the cooler months, especially November (66), December (100), and January (25).

Figures

FIGURE 131. Known western North American localities of Macrohydnobius contortus (black dots).

FIGURES 123–130. Macrohydnobius contortus. 123. Right antenna, dorsal view. 124. Male mandibles, dorsal view. 125. Female mandibles, dorsal view. 126. Male right protibia. 127. Aedeagus, dorsal view. 128. Everted internal sac, dorsal view. 129. Female sternite 8, ventral view. 130. Female coxites and styli, ventral view. Scale bar = 0.3 mm (Fig. 130), 0.5 mm (Figs. 127–129), 0.75 mm (Fig. 123), 1.0 mm (Figs. 124–126).