Cyclocephala borealis Arrow, 1911

Moore, Matthew R., Cave, Ronald D. & Branham, Marc A., 2018, Annotated catalog and bibliography of the cyclocephaline scarab beetles (Coleoptera, Scarabaeidae, Dynastinae, Cyclocephalini), ZooKeys 745, pp. 101-378 : 144-145

Cyclocephala borealis Arrow, 1911

Cyclocephala borealis Arrow, 1911: 172 [original combination, new replacement name for Cyclocephala villosa Burmeister).

syn. Cyclocephala villosa Burmeister 1847: 54 [original combination, homonym of Cyclocephala villosa Blanchard].

Cyclocephala angularis (Knoch) [synonymy by Harold 1869b: 1241].

Cyclocephala villosa Burmeister [revalidated species status by Horn 1871: 336-337].

Ochrosidia (Ochrosidia) villosa (Burmeister) [new combination and new subgeneric classification by Casey 1915: 147].

Cyclocephala borealis Arrow [synonymy by Arrow 1937b: 8].

Types.

Invalid neotype ♂ of C. villosa at HNHM ( Endrődi Collection) ( Endrődi 1966).

Distribution.

UNITED STATES: Alabama, Arkansas, Connecticut, Delaware, District of Columbia, Georgia, Illinois, Indiana, Kansas, Kentucky, Louisiana, Maryland, Michigan, Mississippi, Missouri, New Jersey, New York, North Carolina, Ohio, Pennsylvania, Rhode Island, South Carolina, Tennessee, Virginia, West Virginia.

References.

Knoch 1801, Burmeister 1847, Melsheimer et al. 1853, Reiche 1859, Harold 1869b, Horn 1871, Crotch 1873, Henshaw 1885, Fall 1901, Fall and Cockerell 1907, Blatchley 1910, Swenk 1911, 1913, King 1914, Casey 1915, Hayes 1918, Arrow 1911, 1937b, Leng 1920, Dawson 1922, Leonard 1923, Sim 1934, Anderson 1936, Britton 1932, 1938, Metcalf and Flint 1939, Sanderson 1940, Johnson 1941a, b, Blackwelder 1939, Riegel 1942a, 1948, White 1947, Blackwelder and Blackwelder 1948, Fleming 1948, Adams 1949, Metcalf et al. 1951, Neiswander 1938a, b, 1951, Peterson 1951, Jaques 1951, Bigger and Blanchard 1955, Maddock and Fehn 1958, Harris 1959, Martínez 1954, Dutky 1941, 1963, Polivka 1950, 1952, 1959, 1960, 1965, Howden and Endrődi 1966, Ritcher 1958, 1966, App and Kerr 1969, Beard 1972, Swan and Papp 1972, Woodruff 1973, Gerhardt and Stanghellini 1971, Kawanishi 1974, Pike et al. 1976, MacLean 1977, Pike et al. 1977, Meyer 1978, 1979, 1981, Potter 1980, 1981, 1995, 1998, Warren and Potter 1983, Bulla et al. 1985, Endrődi 1966, 1985a, Tashiro 1987, Klein 1988, Sutton 1988, Gaugler and Georgis 1991, Hardy 1991, McNamara 1991, Youngman et al. 1993, Suggars Downing 1994, Haynes and Potter 1995, Poole and Gentili 1996, Potter et al. 1996, Fuxa 1998, Peck and Thomas 1998, Bhargava 1999, Brandenburg and Royals 1999, Rothwell and Smitley1999, Vittum et al. 1999, Held et al. 2000a, b, c, Harpootlian 2001, DeFoliart 2002, Cappaert and Koppenhöfer 2003, Riley and Wolfe 2003, Rogers et al. 2003, Wilson et al. 2003, Cranshaw 2004, Grewal et al. 2001, 2002, 2004, Rogers and Potter 2004a, b, c, Koppenhöfer and Grewal 2005, Grewal et al. 2005, Shapiro-Ilan and Cottrell 2005, Harris 2006, Georgis et al. 2006, Jackson and Klein 2006, Heller and Walker 1999a, b, c, d, 2000a, b, 2001a, b, c, d, 2002a, b, c, d, e, f, g, 2003a, b, c, d, e, f, g, 2004a, b, c, 2005a, b, c, d, e, Heller and Kline 2005a, b, c, 2007a, b, c, d, e, f, g, h, i, Bixby et al. 2007, An and Grewal 2007, Ratcliffe and Paulsen 2008, Dingman 2008, Koppenhöfer et al. 2002b, 2005, 2006, 2007, 2008, 2013, Counts and Hasiotis 2009, Koppenhöfer and Fuzy 2003a, b, 2006, 2008a, b 2009, Krinsky 2009, Heller et al. 2000a, b, 2006a, b, c, d, e, f, 2008a, b, c, d, e, f, g, h, i, j, k, l, m, 2009a, b, c, d, e, f, g, h, i, j, k, l, Li et al. 2007, 2009, Peck 2009, Popay 2009, Power et al. 2009, Samples et al. 2009, Smith 2003, 2009, Bélair et al. 2010, Ennis et al. 2010, Holmstrup et al. 2010, Mashtoly et al. 2009, 2010, Redmond and Potter 2010, Jordan et al. 2012, Krajcik 2005, 2012, Redmond et al. 2012a, b, Bousquet et al. 2013, Gardner 2013, Guo et al. 2013, Ranger et al. 2009, 2013, Williamson et al. 2004, 2008, 2013, Hussaini 2014, Anderson et al. 2015, Behle et al. 2015, Chong and Hinson 2015, Renkema et al. 2015, Shetlar and Andon 2013, 2015, An and Grewal 2016, Behle and Goett 2016, Christians et al. 2016, Dossey et al. 2016, Gyawaly et al. 2016, Koppenhöfer and Wu 2016, Mitasuhashi 2016, Park and Tak 2016, Schrader et al. 2016, Subramanian and Muthulakshmi 2016, Del Valle et al. 2017, Ratcliffe and Cave 2017.

Remarks.

A specimen of C. borealis was reported from Mexico (Durango), but this record is considered erroneous ( Endrődi 1966, 1985a, Ratcliffe and Morón 1997, Ratcliffe et al. 2013). Records for Iowa, Nebraska, and Texas for C. borealis are doubtful or spurious ( Ratcliffe and Cave 2017). Cyclocephala borealis has been recorded in Canada (Ontario and Nova Scotia) ( McNamara 1991, Bousquet et al. 2013), though major faunistic studies did not report additional data ( Ratcliffe and Cave 2017).