Adapsiliina Rondani, 1869

Korneyev, V. A., 2016, New Taxa And Synonymy In The Family Pyrgotidae (Diptera Tephritoidea) Ii Subtribe Adapsiliina And Afrotropical Campylocera, Vestnik Zoologii 50 (3), pp. 195-218 : 196

Subtribe Adapsiliina Rondani, 1869 , revised rank

Adapsilioidi Rondani, 1869: 6; Sabrosky, 1999: 39.

Type genus: Adapsilia Waga, 1842 .

D i a g n o s i s. Ocelli always absent; frontal setae lacking; antennae usually moderately or very long, commonly with pedicel and flagellomere more than 1.5 × as long as wide, pedicel without incision (A); face with long, separate or fused antennal grooves; epistome low, sclerotized (A); wing with subcostal vein entire and reaching costal vein at acute angle (?A); costal vein without humeral break (A) but often with inconspicuous subcostal break (A); abdomen with sternites 1 and 2 fused (SA with other Pyrgotinae ) without a seam (A); male genitalia: phallic guide well developed (A); epandrium bare dorsally (A); inner surstylus without prensisetae (A); female: oviscape on ventral side without tongue-like combs of thickened setae or unpaired lobe (A? as reversal); eversible membrane without taeniae (A); aculeus moderately small, at most 1.5× as long as oviscape width at aperture (A), with sternite 8 lacking setae (A) and no anal slit (A); ventral receptacle scepter-like (SA with other Pyrgotinae ). In addition, many of the members of this subtribe possess the so-called femoral organ (an area with thin cuticle devoid of setae) on anterior surface of the mid femur, the character not occurring anywhere outside Adapsiliina (only a few Australian Toxura (Toxurini) and South American Tropidothrinax (Pyrgotina) have somewhat similar structures on fore femur and fore tibia).

Monophyly of this subtribe is supported by at least one synapomorphy: the presence of subocular sclerite (secondary sclerotised crescent area on the gena ventral of the compound eye) and epandrium without setae on its dorsal side. Its representatives have several characters, which are believed to be synapomorphies of the subfamily Pyrgotinae (abdominal sternites 1 and 2 fused, ventral receptacle scepter-like), Pyrgotini +Toxurini (prensisetae lacking), and Pyrgotini (subcosta straight, without subapical bent). Distribution of the character modalities among other Pyrgotini and Toxurini remains very poorly examined, and even the possible sister-group of the Adapsiliina is unknown.

Th e following Old World genera belong here:

Adapsilia Waga, 1842 ; Campylocera Macquart, 1843 ; Euphya Wulp, 1885 ; Eupyrgota Coquillett, 1899 ; Geloemyia Hendel, 1908 ; Hendelpyrgota Vanschuytbroek 1963 ; Plectrobrachys Enderlein, 1942; Porpomastix Enderlein, 1942 ; Pyrgotomyia Hendel, 1934 ; Siridapha Enderlein, 1942 ; Trichopeltia Wulp, 1885 . In addition, a group of species related to Adapsilia hirtoscutellata Hendel, 1933 forming an unnamed and unplaced genus-group taxon, belongs in this subtribe.