Colocistis thermarum (Bradley)

Kimsey, Lynn S. & Wasbauer, Marius S., 2013, Revision of the brachycistidine genus Colocistis Krombein, 1942 (Hymenoptera, Tiphiidae), Journal of Hymenoptera Research 33, pp. 1-24 : 18-19

Colocistis thermarum (Bradley) Figures 8 , 18 , 30 , 40 , 48 , 56

Brachycistis thermarum Bradley, 1917:274. Holotype male; USA: Arizona, Hot Springs (CUIC).

Diagnosis.

This is one of three species, including Colocistis castanea and Colocistis oaxacana , which have three submarginal cells and the second metasomal sternum simple. It can be distinguished from both by the tuberculate clypeus.

Description. Male.

Body length. 8-15 mm.

Head ( Fig. 8 ). Compound eyes enlarged, extending well beyond temples; ocelli enlarged, diameter of lateral ocellus at least 0.7 × OOL; impressed line present between lateral ocelli; antennal socket rim only slightly thickened ventrolaterally; clypeus with median tubercle visible above mandible in lateral view, apex not bounded laterally by strong teeth; gular carina not raised; LID 0.7 × UID; flagellomeres I-II length 2.3-2.4 × breadth; flagellomere IX length 5 × breadth; occipital carina strong but, complete ventrally; hypostomal carina not raised; mandible narrowest submedially, apically broad and teeth subequal in length.

Mesosoma. Pronotum nearly vertical anteriorly, punctures deep, 0.5 puncture diameter apart to contiguous, shoulders well-developed, not produced laterally; forefemur without stout spines; foretibia without spines except apically; basitarsus with two spines on outer surface; tibial spur 0.9 × length of basitarsus; hindcoxa without basally tooth-like longitudinal dorsal carina; scutal punctures 0.5-2.0 puncture diameters apart; mesopleural punctures contiguous; scutellum flattened dorsally nearly planar with level of propodeum, punctures 0.5-1.0 puncture diameters apart, with small, polished impunctate area medially; postscutellum densely punctate; propodeal dorsal surface separated from posterior and lateral surfaces by carina, dorsal surface with longitudinal medial trough, surface shallowly rugose, posterior surface nearly impunctate, lateral surface densely punctate dorsally, impunctate ventrally; forewing with 3 submarginal cells; costa extending along wing beyond stigma.

Metasoma. Tergum I without transverse carina; sternum II without raised area basally.

Genitalia ( Figs 48 , 56 ). gonostylar apex nearly straight in lateral view; aedeagus abruptly decurved before apex, without dorsally projecting process.

Color. Head and body unicolorous, castaneous; vestiture whitish to pale yellow.

Female unknown.

Distribution

( Fig. 40 ): USA: Arizona: Cochise, Maricopa, Pima, Pinal, Yuma counties; California: Imperial, Inyo, Riverside counties; Texas: Culberson, Dimmit, Jeff Davis counties; MEXICO: Baja California, Baja California Sur, Sonora; 370 males were examined (BME, CAS, CDFA, EMEC).

Figures

Figures 1 - 12. 1 - 8 Front view of male head. 9 Lateral view of male head 10 - 12 Ventral view of female head. Abbreviations: LID = least interocular distance, OcC = occipital carina; OOL = ocellocular distance, THC = transverse hypostomal carina, UID = upper interocular distance.

Figures 13 - 24. 13, 14 Male forewing. 15 Ventral view of male hindcoxa 16 Dorsal view of male propodeum 17 dorsal view of male mandible 18 lateral view of mandible 19 Ventral view of male metasomal sterna I-II 20, 22 Lateral view of male metasomal segments I and II 21, 23, 24 Ventral view of female metasomal sternum I. Abbreviations: SM- 1, 2, 3 = submarginal cells 1 - 3.

Figures 25 - 32. 25 - 28. Males. 29 - 32 Females. 25, 27, 29, 31 Lateral view 26, 28, 30, 32 Dorsal view 25 - 30 are images of the holotypes.

Figures 37 - 40. Distribution maps Colocistis species.

Figures 41 - 48. Ventral view of male genital capsule.

Figures 49 - 56. Lateral view of male genital capsule.