Trogoderma sahondrae, Háva & Baňař, 2017

Háva, Jiří & Baňař, Petr, 2017, A new TrogodErma species from Central Madagascar (Coleoptera: Dermestidae: Megatominae), Zootaxa 4299 (2), pp. 279-284 : 280-283

Trogoderma sahondrae sp. nov.

( Figs. 1–10 )

Type material. Holotype, male: ABT \ Nov.2011 \19 MADAGASCAR / AMBOHITANTELY Spec. Res. ; 1497m / S18°10´56.9´´E47°17´21.9´´; 23.xi. / 2011; sifting forest litter; Winkler app. / extraction; L.S. Rahanitriniaina lgt. (MMBC). Paratypes: (1 spec.) the same data as holotype ( MMBC) ; (9 spec.): ABT \ Nov.2011 \15 MADAGASCAR / AMBOHITANTELY Spec. Res. ; 1634m / S18°11´32.0´´E47°17´08.7´´; 21.xi. / 2011; sifting forest litter under / Pandanus ; Winkler app. extraction; / L.S. Rahanitriniaina lgt. (4 MMBC, 5 JHAC); (4 spec.): ‘ ABT \ Nov.2011 \02 MADAGASCAR / AMBOHITANTELY Spec. Res. ; / S18°11´49.1´´ E47°17´05.6´´ ; 1547m / 16–17.xi.2011; sifting forest litter / Winkler app. extraction / L.S. Rahanitriniaina & P. Baňař lgt. ’ (2 MMBC, 2 JHAC); (3 spec.): ‘ ABT \ Nov.2011 \14 MADAGASCAR / AMBOHITANTELY Spec. Res. ; 1623m / S18°11´22.7´´E47°17´07.8´´; 21.xi. / 2011; sifting forest litter under / Pandanus ; Winkler app. extraction; / L.S. Rahanitriniaina lgt. ’ (2 MMBC, 1 JHAC); (7 spec.) ABT \ Nov.2011 \07 MADAGASCAR / AMBOHITANTELY Spec. Res. ; 17.xi. / 2011; S18°11´48.5´´ E47°17´09.3´´ / sifting forest litter under Pandanus / Winkler app. extraction; very dry / material ; 1595m; P. Baňař lgt. (4 MMBC, 3 JHAC); (1 spec.) ABT \ Nov.2011 \16 MADAGASCAR / AMBOHITANTELY Spec. Res. ; 1620m / S18°11´42.6´´E47°17´10.8´´; 21.xi. / 2011; sifting forest litter under / Pandanus ; Winkler app. extraction; / L.S. Rahanitriniaina lgt. (MMBC); (1 spec.): ‘ ABT \ Nov.2011 \10 MADAGASCAR / AMBOHITANTELY Spec. Res.; 18.xi. / 2011; sifting forest litter under palm / tree; Winkler app. extraction; / ~ 1600m; P. Baňař lgt. ’ ( MMBC). [most of the paratypes card mounted, some of them dissected or damaged, those alcohol-preserved].

Description. Male. Body measurrements (in mm, taken from four specimens):

Total body length: 2.55–2.81. Head: total length: 0.48–0.50; maximum width across eyes: 0.69–0.71; minimum distance between eyes: 0.40–0.41. Pronotum: maximum (median) length: 0.69–0.71; maximum width: 1.47–1.52. Elytra: maximum length: 2.03–2.18; maximum width: 1.65–1.72.

General appearance. Small, oval species with conspicuous colour pattern ( Figs. 1–3 ).

Colouration. Dark-brown on dorsal and ventral surfaces ( Figs. 1–3 ), legs paler, light brown, scape, pedicel and antennomere IX light brown, antennomeres III–VIII yellowish, antennomeres X and XI dark brown to blackish.

Structure. Head finely punctate with regularly distributed short black setae. Palpi entirely yellowish-brown; setation on mentum dense. Eyes small, not overlapping lateral margin of head from dorsal view, ocular index 2.58. Ocellus on frons conspicuous. Antennae ( Fig. 4 ) composed of 11 antennomeres, antennal club of 4 antennomeres. Thorax. Pronotum finely punctate like head with short black setation; lateral and anterior margins continuous, without any crank or angle regularly rounded, posterior margin conspicuously bisinuate, anterior angles not visible from above. Pronotum much wider than long ( Fig. 1 ), pronotum width to maximum length ratio 2.13. Mesonotum heavily sclerotized ( Fig. 6 ), scutellum small, triangular, finely punctate as for pronotum and head, asetose. Metanotum robust ( Fig. 6 ), more weakly sclerotized than mesonotum, hind wing as in Fig. 7 . Meso-metasternum dark brown with short golden setation.

Elytra coarsely punctate; brown with three, transverse, orange fasciae ( Fig. 5 ); covered by golden setation ( Figs. 1–3 ). Anterior half of elytral epipleuron conspicuously darker than rest of elytron, entirely brown to blackish. Ratio of elytra length to their maximum width 1.27. Legs brown with short golden setation; tibiae without spines. Abdominal ventrites with short golden setation ( Fig. 2 ). First visible abdominal ventrite with distinct oblique discal striae and robust, heavily sclerotized median tubercle present ( Fig. 8 ). Pygidium brown with short, black setation. Male genitalia. Aedeagus narrow, parallel-sided in apical two thirds ( Figs. 9–10 ), heavily sclerotized basally.

Variability. Total body length 2.55–2.81 mm, maximum elytral width 1.65–1.72.

Sexual dimorphism. Female similar to male, but terminal antennomere conspicuously smaller.

Differential diagnosis (Madagascan species only included). The new species differs from T. taomasinum Háva, 2009 by bicolorous elytra (unicolorous in T. taomasinum ); from T. trifasciatum Háva, 2009 by conspicuously oval body shape (body elongate in T. trifasciatum ); from T. fasciolata (Fairmaire, 1897) , T. impressiceps (Pic, 1915) , T. madecassum (Pic, 1924) , T. wolfgangi Háva & Herrmann, 2008 and T. housei Háva, 2014 by antennal club composed of four antennomeres (antennal club composed of three antennomeres in T. fasciolata and T. impressiceps ; five antennomeres in T. wolfgangi and T. housei and six antennomeres in T. madecassum ); from T. seminigrum Pic, 1915 by presence of large orange fasciae on elytra (small spots of whitish pubescence in T. seminigrum ) and from T. horaki Háva, 2013 and T. sambiranum Háva, 2009 by presence of three large orange fascia on elytra (one orange fascia in T. horaki , one orange spot on apex of each elytron and single orange fascia of white pubescence in T. sambiranum ). See also the Identification key to the known Madagascan species of Trogoderma following.

Etymology. This species is named after Lalao Sahondra Rahanitriniaina, our Malagasy colleague who collected most of the type series.

Collecting circumstances. All known specimens were sifted from leaf litter in mountain forests of Ambohitantely Special Reserve. Sifted material was subsequently extracted in a Winkler apparatus and collected into alcohol.

Type locality. Ambohitantely Special Reserve , Central Madagascar.

Figures

FIGURES 1 – 3. Trogoderma sahondrae Háva & Baňař sp. nov., male paratype. 1, dorsal view; 2, ventral view; 3, lateral view. Scale bar in mm.

FIGURES 4 – 8. Trogoderma sahondrae Háva & Baňař sp. nov., male paratype. 4, left antenna; 5, left elytron; 6, meso- and metanotum; 7, right hind wing; 8, abdominal sternites, ventral view. Scale bars in mm.

FIGURES 9 – 10. Trogoderma sahondrae Háva & Baňař sp. nov., aedeagus of male paratype. 9, ventral view; 10, dorsal view. Scale bars in mm.