Neobisium (Neobisium) herzegovinense Beier, 1939

Novák, János & Hörweg, Christoph, 2017, Redescriptions Of Three Species Of The Genus Neobisium From The Balkans And The Carpathians (Arachnida: Pseudoscorpiones), Acta Zoologica Academiae Scientiarum Hungaricae 63 (3), pp. 277-292 : 289-290

Neobisium (Neobisium) herzegovinense Beier, 1939

(Figs 4G–J)

Typematerialexamined (syntypes): 1 m, 2 f, BosniaandHerzegovina, fromhigher areaoftheVolujakMts, understones, 2114 masl., 09.07.1912 (NHMW-25216).

Description. Medium-sizedpseudoscorpionwithepigeanhabitus. Carapace, cheli- ceraeandpedipalpsyellowishbrown, opisthosomaandlegsyellowish. Measurementsand ratiosofthestudiedspecimensasinTable 3.

Carapaceapproximatelyaslongasbroad, surfacesmooth (Fig. 4G). Epistometri- angularandprominent. Twopairofeyeswithlens. Onepairofslitlikelyrifissuresatan- teriorandonepairatposteriorhalfofcarapace. Preocularsetaeabsent. Setalformula: 4:6:6–8:11–12 (27–30).

Coxalarea. Manducatoryprocesswith 5–6 setae, restofpedipalpalcoxawith 8–14 se- tae; pedalcoxaIwith 8–13 setae; II: 9–12; III: 8–11; IV: 13–17. Anterolateralprocessofcoxa Ilong, triangularandapicallypointed; medialprocessroundandflattened, withdenticles.

Opisthosomawithgranulostriatepleuralmembrane. ChaetotaxyoftergitesI–X: 8–10:12–14:14–16:16:16–20:16–20:16–18:16–18:16–18:10–12. SegmentXIwith 16–18 setae, 4–6 ofthemtactile. Analconewith 2 ventraland 2 dorsalsetae. Malegenitalregion: sternite II with 14 small setae, sternite III with 27 setae, of which 13 along posterior margin. Numberofinternalsetaeinmaleopeningandthemedialgenitalsackcouldnotbeinvestigated. Female genital region: sternite II with 10–13 small setae, sternite III with 17–21 small setae in a row. Chaetotaxy of sternites IV–X: 16–24:20–26:20–28:20–27:20–28:19–23:16–18.

Chelicerawith 7 setaeonhand (Fig. 4H). Subgalealseta 0.62–0.69× frombaseofmov- ablefinger. Spinneretaflattenedhyalinetuberclewith 9–13 silkducts. Fixedfingerwith 9–16 smallandmedium-sizedteeth, movablefingerwith 8–10 smallandmedium-sized teeth. Rallumwith 8 blades, twodistalmostbladesunilaterallypinnateonanteriorface. Thedistalmostbladewithenlargedbasisandseparatedfromtheproceedingone. Serrula exterior with 24–27 blades, serrula interior with 25–27 blades.

Pedipalp (Fig. 4I) surfacessmooth. Trochanterwithtubercle. Femur 4.22–4.60, patella 2.65–2.96 timesaslongasbroad. Patellawith 2 lyrifissuresonbasalpartand 2 smaller nearthearticulationwiththehand. Pedicelofpatellabears 2–4 micropores. Chela (Fig. 4J). Pedicelbears 2–4 dorsalmicropores. Chelalfingersapproximatelyaslongashandwith pedicel. Fixedchelalfingerwith 66–71 close- setandpointedteethofequallength. Movable chelalfingerwith 57–68 roundedteeth, dentallineendingattheleveloftrichobothrium b. TrichobothriaofchelalfingersdistributedasinFigure 4J. Distancebetweentrichobothria ib and ist approximatelyaslongasbetween ist and it.

Legswithsmoothsurfaces. Clawssmoothandsimple, aroliashorterthanclawson alllegs. SubterminalsetaoftelotarsusIVwithtwoserraterami. Tactilesetaepresenton tibia, basitarsusandtelotarsusoflegIV.

Remarks. ThespecieswasdesribedbyBeieras Neobisium (Neobisium) herzegovinense fromonemaleandtwofemaletypes, fromtheVolujakMts (BosniaandHerzegovina) ( BEIER 1939 b). LaterBEIER (1963) mentioned N. herzegovinense as a subspecies of N. bosnicum as N. bosnicum herzegovinense . Since itsdescriptionnofurtheroccurrencesofthespecieswerepublished.

Duringthepresentinvestigationwefoundgreatervariabilityinthesetationofthecarapaceandthetergites, andinthemeasurementdataandpro- portionsthanpreviouslymentionedintheliterature ( BEIER 1939 b, 1963).

Consideringtheimportantdifferencesbetween N. (N.) bosnicumbosnicum and N. (N.) bosnicumherzegovinense inthemeasurements, ratiosandshapeof thepedipalps, itiswarrantedtore- elevate N. (N.) bosnicumherzegovinense to fullspeciesrank.