Amplirhagada albertiana, Köhler & Johnson, 2012

Köhler, Frank & Johnson, Michael S., 2012, Species limits in molecular phylogenies: a cautionary tale from Australian land snails (Camaenidae: Amplirhagada Iredale, 1933), Zoological Journal of the Linnean Society (Zool. J. Linn. Soc.) 165 (2), pp. 337-362 : 356-358

publication ID	https://doi.org/ 10.1111/j.1096-3642.2011.00810.x
DOI	https://doi.org/10.5281/zenodo.10543004
persistent identifier	https://treatment.plazi.org/id/D2543F71-FC26-FF9B-B8E2-FCBE5CC8C0BC
treatment provided by	Marcus
scientific name	Amplirhagada albertiana
status	sp. nov.

Treatment

AMPLIRHAGADA ALBERTIANA View in CoL SP. NOV.

FIGURES 7V View Figure 7 , 18 View Figure 18 , 19 View Figure 19

Type locality: Western Australia, Kimberley, Bonaparte Archipelago, Albert Island , 14°31′09.1″S, 124°55′48.4″E GoogleMaps .

Material examined: Holotype WAM S42947 View Materials (dissected specimen) . Paratypes WAM S33000 View Materials (four preserved specimens) AM C.470992 (one preserved body) ( Table 1). Additional, non-type material . FMNH 219169-70 About FMNH , FMNH 219670 About FMNH from Albert Island (KC-043, coll. 14.vii.1988, V. Kessner) .

Etymology: For Albert Island, adjective of feminine gender.

Description: Shell ( Fig. 7V View Figure 7 ) globose to broadly conical with medium high spire, thick, periphery evenly rounded ( Table 3); background and ventral whitish colour crème, with brown, sub-sutural, and mid-whorl bands, shell tip may be covered in brownish suffusion; outer lip white; teleoconch with axial growth lines only. Umbilicus 50–100% concealed by columellar reflection ( Table 4). Aperture simple, slightly to well expanded, slightly reflected, with weak or wanting basal node. Genital anatomy ( Figs 18 View Figure 18 , 19 View Figure 19 ). Penis inflated, long, coiled within penial sheath; vas deferens entering penial sheath within anterior third, penial retractor muscle moderate in length; penial verge short (<1/10 length of penial chamber), pointed; inner penial wall without main pilaster; posterior third of wall with densely packed, elongate, roundish, irregularly arranged pustules; anterior half of inner wall with irregularly shaped and arranged, corrugated, indistinct, oblique pilasters. Vagina long (about same length as penis), tubular; bursa copulatrix moderately long, tubular, extending to anterior end of spermoviduct, with inflated head.

Comparative remarks: Shell most similar to A. berthierana in shape and colour but smaller (similar to and A. turbinensis ), differing from the former two species by closed instead of open umbilicus. Unlike A. intermedia , A. berthierana , and A. turbinensis penial wall not entirely covered with pustules. Typical features are coiled penis and absence of main pilaster. Bursa copulatrix conspicuously elongated (even more so than in former species). Material kept in the FMNH is labelled as ‘ Amplirhagada sp 69’.

Distribution: Albert Island only ( Fig. 6 View Figure 6 ).

AMPLIRHAGADA GEMINA KÖHLER, 2010

FIGURE 20A–D View Figure 20

Amplirhagada gemina Köhler, 2010a: 270 View in CoL View Cited Treatment , figs 82–84.

Type locality: Western Australia, Kimberley mainland near Camden Sound, 4.5 km north of Prior Pt , 1.5 km south of Hall Pt , 15°40′45″S, 124°22′45″E GoogleMaps .

Material examined ( Table 1): Holotype WAM S34611 View Materials ( Fig. 20A View Figure 20 ) . Paratypes AM C.463754, WAM S33029 View Materials . Non-type material. WAM S42955 View Materials -7 About WAM ( Fig. 20B–D View Figure 20 ).

Diagnosis: Shell ( Fig. 20A–D View Figure 20 ) semi-globose to broadly conical, with moderately elevated spire; periphery evenly rounded to slightly angulate; with regular growth lines only. Umbilicus 60–100% concealed by columellar reflection ( Table 4). Colour brown, base beige, lip colour whitish. Aperture simple, slightly expanded, weakly reflected, basal node absent. Genital anatomy. Penis slightly inflated, moderately long, straight; vas deferens entering penial sheath apically, penial retractor muscle moderate in length; penial verge tiny; inner penial wall entirely covered with narrowly packed, elongate, rather small pustules, no main pilaster differentiated. Vagina short (shorter than penis), inflated; bursa copulatrix moderately long, wide, extending to anterior end of spermoviduct, with weakly inflated head ( Köhler, 2010a). Comparative remarks: Amplirhagada gemina and A. kimberleyana are difficult to differentiate. Their shells look almost identical, with specimens from Lulim Island being more similar to A. kimberleyana than to conspecific specimens from the mainland ( Fig. 3 View Figure 3 ). Main differentiating feature is slightly different penial wall sculpture: A. gemina has more crowded, smaller, undifferentiated pustules while A. kimberleyana has larger and fewer pustules, arranged in rows, some of them with hooks forming a weak main pilaster.

Distribution: South-west Kimberley coast between south of Halls Pt and Wilsons Pt, including Lulim Island, close to mainland coast ( Fig. 6 View Figure 6 ).

AMPLIRHAGADA KIMBERLEYANA KÖHLER, 2010

FIGURE 20E–F View Figure 20

Amplirhagada kimberleyana Köhler, 2010a: 268–9 View in CoL View Cited Treatment , figs 79–81.

Type locality: Western Australia, northern Slate Island, Camden Sound, 2 km north-west of Wilsons Pt , 15°32′10″S, 124°23′56″E GoogleMaps .

Material examined ( Table 1): Holotype WAM S34611 View Materials ( Fig. 20E View Figure 20 ) . Paratypes AM C.463754, WAM S33029 View Materials . Non-type material. WAM S42955 View Materials -7 About WAM ( Fig. 20F View Figure 20 ).

Taxonomic remarks: The type material was collected in 1988 on the ‘northern Slate Island’, which is identical to Wailgwin Island, named so subsequently.

Diagnosis: Shell ( Fig. 20E, F View Figure 20 ) semi-globose to broadly conical, with moderately elevated spire; periphery evenly rounded to slightly angulate; with regular growth lines only. Umbilicus 60–100% concealed by columellar reflection ( Table 4). Colour brown, occasionally banded, base and background colour beige, lip colour whitish. Aperture simple, slightly expanded, weakly reflected, basal node absent. Genital anatomy. Penis slightly inflated, moderately long, straight; vas deferens entering penial sheath within anterior third, penial retractor muscle moderate in length; penial verge tiny; inner penial wall entirely covered with narrowly packed, elongate, rather small pustules, main pilaster weakly differentiated, consisting of 2–3 parallel rows of inconspicuously enlarged pustules that support little hooks. Vagina short (shorter than penis), inflated; bursa copulatrix moderately long, wide, extending to anterior end of spermoviduct, with weakly inflated head ( Köhler, 2010a).

Comparative remarks: Material studied here is topotypic. See above for comparison with A. gemina .

Distribution: Wailgwin Island only ( Fig. 6 View Figure 6 ).