Amplirhagada imperialis, Köhler & Johnson, 2012

Köhler, Frank & Johnson, Michael S., 2012, Species limits in molecular phylogenies: a cautionary tale from Australian land snails (Camaenidae: Amplirhagada Iredale, 1933), Zoological Journal of the Linnean Society (Zool. J. Linn. Soc.) 165 (2), pp. 337-362 : 358-359

publication ID	https://doi.org/ 10.1111/j.1096-3642.2011.00810.x
DOI	https://doi.org/10.5281/zenodo.5479892
persistent identifier	https://treatment.plazi.org/id/D2543F71-FC20-FF98-B8EF-FC625B78C031
treatment provided by	Marcus
scientific name	Amplirhagada imperialis
status	sp. nov.

Treatment

AMPLIRHAGADA IMPERIALIS View in CoL SP. NOV.

FIGURES 21A View Figure 21 , 22 View Figure 22 , 23 View Figure 23

Type locality: Western Australia, Kimberley, Bonaparte Archipelago, southern section of Boongaree Island , 15°05′55.8″S, 125°10′27.4″E GoogleMaps .

Material examined ( Table 1): Holotype WAM S42948 View Materials (dissected specimen) . Paratypes WAM S42949 View Materials (two preserved specimens, Fig. 21A View Figure 21 ), AM C.470993 (two preserved bodies) ( Table 1) .

Etymology: For ‘imperialis’ (Latin: ‘of the empire or emperor’), in reference to King Bungaree, name patron of Boongaree Island; adjective of feminine gender.

Description: Shell ( Fig. 21A View Figure 21 ) beehive-shaped, highly turreted with angulated periphery, flattened whorls, shallow suture (H = 18.2 ± 0.6, D = 20.5 ± 0.8, W = 5.4 ± 0.05, average ± SD of three shells); background and ventral whitish crème, with diffuse brown, subsutural and mid-whorl bands, usually blending into each other covering shell in brownish suffusion; outer lip light purplish; teleoconch with regular axial growth lines. Umbilicus completely concealed by columellar reflection. Aperture simple, slightly expanded, slightly reflected, with weak basal node. Genital anatomy ( Figs 22 View Figure 22 , 23 View Figure 23 ). Penis inflated, long, straight; vas deferens entering penial sheath apically, penial retractor muscle moderate in length; penial verge short (<1/6 length of penial chamber), elongated; inner penial wall with large, conical, corrugated main pilaster, comprising entire length of penial wall, broadest at posterior end, tapering in thickness and development anteriorly, one or two additional pilasters of similar structure forming along entire length of inner penial wall being reduced in prominence in thickness; inner penial wall entirely covered by flat, indistinct, partly fused pustules. Vagina long (about same length as penis), tubular; bursa copulatrix moderately long, tubular, extending to anterior end of spermoviduct, with inflated head.

Comparative remarks: From all species described herein, A. imperialis clearly differs in its beehiveshaped, highly turreted shell. Three congeneric species have been described from the same island, of which A. boongareensis is most similar ( Fig. 21B View Figure 21 ). However, it is even larger and more strongly elevated, having more strongly developed growth lines. Amplirhagada boongareensis differs most conspicuously in its inner penial wall sculpture, consisting of densely packed, small pustules that cover the entire surface of the penial wall as well as having a huge main pilaster. The present species may be conspecific with unidentified shells from the central section of Boongaree Island reported by Köhler (2010a) as Amplirhagada sp. , in which case all known species on this island occurred in sympatry.

Distribution: Boongaree Island only ( Fig. 6 View Figure 6 ).