Aysenia barrigai, Izquierdo, Matías A. & Ramírez, Martín J., 2008

Izquierdo, Matías A. & Ramírez, Martín J., 2008, Two new spider species of the genera Aysenia and Aysenoides from southern Chile and Argentina: description and phylogenetic relationships (Araneae: Anyphaenidae, Amaurobioidinae), Zootaxa 1861, pp. 29-43 : 33-37

publication ID	https://doi.org/ 10.5281/zenodo.183660
DOI	https://doi.org/10.5281/zenodo.5659822
persistent identifier	https://treatment.plazi.org/id/D2297B6E-181B-B720-FF04-FCDCFF5D4575
treatment provided by	Plazi
scientific name	Aysenia barrigai
status	sp. nov.

Treatment

Aysenia barrigai View in CoL new species.

Figs 2–11 View FIGURES 2 – 7 View FIGURES 8 – 11

Types: Male holotype and 1 female paratype from Chile, Region IX (Araucanía), Prov. Malleco, Nahuelbuta National Park, S 37º47’, W 73º00’, elev. 1200 m, 12 February 2005, leg. J.E. Barriga T., fogging Nothofagus antarctica , in MHNS.

Additional material examined: ARGENTINA: Río Negro: Nahuel Huapi National Park, Isla Victoria, April. 1945, leg. Havrylenko, 2 females and 2 immatures ( MLP); Chubut: Los Alerces National Park, Lago Futalaufquen, 2 Km from Villa Futalaufquen, 9 February 1986, leg. M. Ramírez, beating bushes in forest of Nothofagus dombeyi , 1 female (MACNAr 13430); Los Alerces National Park, 15–20 March 1974, leg. Bordón, 1 female (MACNAr 13467). CHILE: Región IX (Araucanía): Prov. Malleco, Nahuelbuta National Park, S 37º47’ W 73º00’, elev. 1200 m, 12 February 2005, leg. J. E. Barriga T., fogging Nothofagus antarctica 3 males, 1 female and 3 immatures (MACNAr 13432); same data, fogging Nothofagus dombeyi , 2 males, 2 females and 1 immature (MACNAr 13433).

Etymology: The specific name is a patronym in honor of the collector of the type specimens, Juan Enrique Barriga Tuñón; noun (name) in genitive case.

Diagnosis. Males and females resemble Aysenia segestrioides Ramírez 2003 by the general morphology and body coloration pattern (see Ramírez 2003, fig. 23 A) but can be distinguished by the shape and number of cusps of the paramedian apophysis in the male copulatory bulb ( Figs 8–9 View FIGURES 8 – 11 ), more developed than in other species, while the female vulva has moderately long copulatory ducts and irregular spermathecae, apparently with two or three internal chambers ( Figs 10–11 View FIGURES 8 – 11 ).

Description. Male (holotype): Prosoma length 2.30, width 1.30, more slender than in the female. Length of tibia/metatarsus: I, 1.30/1.25; II, 1.33/1.18; III, 0.93/1.00; IV, 1.33/1.28. Chelicerae smaller, less robust than in the female, with two teeth on retromargin, three on promargin. Length of sternum 1.23, width 0.75. Spines: leg I, femur d 111, p 1ap; tibia v 22 2 (xp 1x slightly to prolateral), p 1d110; metatarsus v 2 02 (r1xr1 slightly advanced and p1xx slightly to prolateral), p 010. II, femur = I; tibia v r1r12, p 0d11; metatarsus v 2 r11 (r1xx slightly advanced), p 01d 1. III, femur d 111, p and r 1ap; tibia d 220 (p1p 1 x slightly advanced) or p and r 110, v 0r12; metatarsus v 2 r12 (r1xx slightly advanced) or v 2 02, p 1ap or p 011, r 011 or r 1ap, d 0p1 2. IV, femur d 111, r 1ap; tibia v p1p12, r 0 d11; metatarsus v 2 p12 or v 2 p1r1p12, p 011, r 011 or r 1ap, d r1r12. Opisthosoma length 2.65, spiracle–epigastrium 1.25, spiracle–spinnerets 0.22.

Color in ethanol ( Figs 6–7 View FIGURES 2 – 7 ): Cephalic region uniform brownreddish margins, ocular area dark, reddishbrown. Chelicerae dark reddishbrown, cheliceral boss paler. Legs brownyellowish with femur and tibia slightly darker. Labium and gnathocoxae reddishbrown, sternum light brown. Opisthosoma brown with cream pattern, spinnerets white. Venter pale, light cream.

Palp: Tibia short, width/length 0.69, RTA long, acute, wider at base. Cymbial conductor wide. Tegulum basal. Sperm duct with two conspicuous loops on anteriordorsal margin. Embolus short. Median apophysis apical, hookshaped. Primary conductor with canal, heavily sclerotized in apical region. Paramedian apophysis with three cusps, two of them heavily sclerotized with common base, closer to median apophysis, third cusp close to primary conductor ( Figs 8–9 View FIGURES 8 – 11 ).

Female (paratype): Prosoma length 2.87, width 1.67, wider in front. Length of tibia/metatarsus: I, 1.30/ 0.80; II, 0.97/0.83; III, 0.73/0.70; IV, 1.30/0.98. Length of palpal tarsus 0.63. Chelicerae very strong, with two teeth on retromargin, three on promargin. Length of sternum 1.48, width 0.80. Spines: Leg I, femur d 111, p 1ap; tibia v 22 2 (xp 1x slightly displaced to prolateral); metatarsus v 2 r10; II, femur = I; tibia v r1r12; metatarsus v 2 r12 or v 2 r1p1, p 010; III, femur d 111, p 1ap; tibia v r 1ap, p 0d1, r 0d1; metatarsus v 2 02, p 011, r 1ap, d 2ap. IV, femur d 111; tibia v p1p12; metatarsus v 22 2, p 1ap, d r 1ap. Leg III directed forward. Femur I–III narrow, IV strong.

Color in ethanol ( Figs 2–4 View FIGURES 2 – 7 ): As in male, except central region of dorsal shield of prosoma redorange.

Opisthosoma length 3.99, epigastrium–spiracle 2.73, spiracle–spinnerets 0.30. Epigyne: Lateral lobes slightly sclerotized, their limits with median field well defined ( Fig. 5 View FIGURES 2 – 7 ). Copulatory ducts difficult to distinguish, apparently not coiled. Ducts of accessory bulbs long. Spermathecae with apparently two or three chambers ( Figs 10–11 View FIGURES 8 – 11 ).

Variability: Spines in males: Metatarsus I, v 22 1 (r1r 1x slightly advanced) or v 22 0, p 110. Metatarsus II, v 22 2 (r1r 1x slightly advanced) or v 2 r1p1p1, p 111. Tibia III, v 2 ap. Metatarsus IV, d 2p1 r1r1p1 or d 022, v 22 2, r 111. Spines in females: femur I, r 1ap. Tibia II, p 1ap. III, v 2 ap, r 11, p 1. Metatarsus III, d 0p12, p 1ap. IV, v 2 p12, r 1ap. Some specimens with dorsum of opisthosoma uniformly dark.

Natural history: Aysenia barrigai new species was collected from shrubs and canopy in forests of southern beech species Nothofagus antarctica and N. dombeyi .

Distribution: Known from Chubut and Río Negro provinces in Argentina, and Malleco province, in Chile.