Ommatius ampliatus, Scarbrough, 2002

Sánchez, Pável & Camargo, Alexssandro, 2023, The ampliatus species group of Ommatius Wiedemann, 1821 (Diptera, Asilidae, Ommatiinae) in Peru with the description of four new species, Zootaxa 5352 (4), pp. 501-520 : 515-517

publication ID

https://doi.org/ 10.11646/zootaxa.5352.4.3

publication LSID

lsid:zoobank.org:pub:460F0F1C-9415-48AB-BCAB-875A0005A981

DOI

https://doi.org/10.5281/zenodo.8435237

persistent identifier

https://treatment.plazi.org/id/D20787A1-FFB6-FFEA-FF13-F9BAFA5FC1CD

treatment provided by

Plazi

scientific name

Ommatius ampliatus
status

 

Key to the species of the ampliatus View in CoL group (modified from Scarbrough 2002)

1. Hind femur mostly to entirely brown to black, sometimes basal third to two-thirds yellow to reddish yellow posteriorly (Figs 1–2, 11, 16, 26, 31, 41, 46–47)........................................................................... 2

1’. Hind femur more extensively yellow, posterior usually mostly yellow to brownish yellow or reddish, apex brown to black..................................................................................................... 21

2. Hind femur with all or most anteroventral macrosetae brown or black, 3–4 basal posteroventral macrosetae often yellow... 3

2’. Hind femur with most anteroventral macrosetae yellow or white, 1–3 anteroventral macrosetae and 3–5 posteroventral macrosetae sometimes brown or black ( Figs 2 View FIGURES 1–10 , 47 View FIGURES 46–52 )........................................................... 8

4. Scutellum with 2 marginal macrosetae; female hind femur slender; female S8 with a large shiny spot laterally, median dorsal stripe and narrow margins with dense pruinosity ( Ecuador)............................ O. depressus Scarbrough, 2002 .

4’. Scutellum with 3 marginal macrosetae; female hind femur wider; female S8 entirely sparse pruinose ( Guyana)...................................................................................... O. constrictus Scarbrough, 2002 .

5. Mid femur mostly yellow posteriorly, apex brown or black.................................................... 6

5’. Mid and hind femora brown to black, narrow base sometimes yellow............................................ 7

6. Mid tibia bent preapically, stout macrosetae present; hind tibia with macrosetae often arising from apical tubercles; T9+10 straight at anterior margin; spermatheca oval ( Venezuela).............................. O. bullatus Scarbrough, 2002 .

6’. Mid and hind tibiae normal, without unusually stout macrosetae arising from apical tubercles; T9+10 with a U indentation with lateral small processes at anterior margin; spermatheca rounded ( Brazil).......... O. aubreyi Vieira, Bravo & Rafael, 2011 .

7. Fore femur mostly yellow; dorsal apical third of epandrium with abrupt, deep emargination, crescent-shaped, apex acutely pointed; gonocoxite with 3–4 setae; gonostylus with two apical processes; phallus tubular, strongly curved dorsally, apex well beyond parameral sheath; female S8 with medioapical margin broadly truncate; spermatheca parabolical ( Costa Rica)................................................................................... O. lunatus Scarbrough, 2002 .

7’. Fore femur mostly black; apical third of epandrium narrow, dorsal margin angled posteriorly to pointed apex; gonocoxite with 1 long thick macroseta; gonostylus narrow, acutely angled apically; phallus with apex beyond parameral sheath; female S8 with medioapical margin produced, corners rounded, spermatheca obovate ( Venezuela)............ O. ayalai Scarbrough, 2002 .

8. Fore and mid femora brown to black, narrow base sometimes red or yellow ( Figs 1 View FIGURES 1–10 , 11 View FIGURES 11–15 )............................. 9

8’. Fore and mid femora at least yellow on basal third to half posteriorly ( Figs 16 View FIGURES 16–25 , 26 View FIGURES 26–30 , 31 View FIGURES 31–40 , 41 View FIGURES 41–45 ).......................... 13

9. Epandrium ending in a narrow process, clavate or rod-like apically ( Fig. 5 View FIGURES 1–10 )...................................... 10

9’. Epandrium ending in an angular process, pointed or rounded apically........................................... 11

10. Wing hyaline, costal cell yellow; epandrium enlarged anteriorly and gradually narrowing to a clavate distal process; subepandrial sclerite only slightly pronounced distally, straight dorsally; gonocoxite with 3 long, thin macrosetae; female cercus slightly concave at distal margin ( Colombia).............................................. O. ampliatus Scarbrough, 2002 .

10’. Wing brownish, costal cell dark brown ( Fig. 3 View FIGURES 1–10 ); epandrium rectangular anteriorly and abruptly narrowing to a rod-like distal process ( Fig. 5 View FIGURES 1–10 ); subepandrial sclerite extremely developed distally forming an arc above the terminalia ( Fig. 5 View FIGURES 1–10 ); gonocoxite without macrosetae ( Fig. 8 View FIGURES 1–10 ); female cercus rounded at distal margin ( Fig.12 View FIGURES 11–15 ) ( Peru)............. Ommatius amaru sp. nov.

11. Epandrium triangular apically ( Fig. 50 View FIGURES 46–52 ); gonocoxite with 1 long, unusually thick, macroseta ( Figs 51–53 View FIGURES 46–52 View FIGURES 53–57 ); phallus narrow apically ( Fig. 55 View FIGURES 53–57 ).................................................................................... 12

11’. Epandrium horizontal apically; gonocoxite with only setae, macroseta absent; phallus wide apically ( Mexico, Guatemala).............................................................................. O. gladiatus Scarbrough, 2002 .

12. Males with vein C dilated anteriorly ( Fig. 48 View FIGURES 46–52 ); subepandrial sclerite wide proximally, tapering distally in ventral view ( Fig. 57 View FIGURES 53–57 ); hypandrium with a small finger-like process distally ( Figs 52–53 View FIGURES 46–52 View FIGURES 53–57 ); gonostylus flat-like basally pronounced externally and bent knee-like medially towards a pointed apex ( Figs 52–53 View FIGURES 46–52 View FIGURES 53–57 ); ejaculatory apodeme pronounced proximal dorsally resembling a golf club, rod-like distally; phallus curved down distally ( Fig. 55 View FIGURES 53–57 ) ( Peru)....................... Ommatius yanantin sp. nov.

12’. Males with vein C not dilated anteriorly; subepandrial sclerite subrectangular in ventral view; hypandrium rounded distally, without process; gonostylus rod-like basally, tapering distally, conic shaped in ventral view; ejaculatory apodeme wide proximally, with a dorsal keel distally; phallus straight to body axis ( Ecuador).................. O. nanciae Vieira, 2015 .

13. Palpus mostly to entirely brown setose................................................................... 14

13’. Palpus entirely yellow setose........................................................................... 16

14. Hind femur dark reddish brown to almost black, extreme base yellow ( Fig. 62 View FIGURES 58–65 ); hind tibia with apical third to apical half dark yellow to brown ( Fig. 58 View FIGURES 58–65 ).............................................................................. 15

14’. Hind femur shining black, extreme base dark reddish; hind tibia with apical half dark brown to almost black ( Fig. 16 View FIGURES 16–25 ) ( Peru)................................................................................ Ommatius kuntur sp. nov.

15. Hind femur dark with most ventral macrosetae yellow ( Fig. 62 View FIGURES 58–65 ); mid femur with only long, posteroventral setae; T9+10 with apical corner concave, spoon-shaped ( Figs 64–65 View FIGURES 58–65 ); S8 with deep notch laterally, surface concave with 1 stout macroseta, corner and medioapical margin equally produced forward ( Figs 64–65 View FIGURES 58–65 ) ( Peru).................. O. quadratus Scarbrough, 2002 .

15’. Hind femur with most ventral macrosetae brown; mid femur with 1 brown, posteroventral macroseta on apical third; T9+10 with apical corner truncate; S8 with medioapical margin far forward of corners, lateral notch with stout macroseta absent ( Peru)........................................................................ O. ovatus Scarbrough, 2002 .

17. Fore and mid femora posteriorly brown on apical half....................................................... 18

17’. Fore and mid femora posteriorly brown on apical fourth to third............................................... 19

18. Scutellum with two brown to black, marginal macrosetae; hind femur usually with mostly yellowish ventral macrosetae, apical 1–2 macrosetae in each row black; long gonocoxal macrosetae present; gonostylus with apical third abruptly angular; hypoproct with basal process slightly produced, J-shaped ( Colombia)........................... O. angulosus Scarbrough, 2002 .

18’. Scutellum with two yellow marginal macrosetae; hind femur with yellow basal macrosetae, apical 5 posteroventral macrosetae black; short, curved gonocoxal macrosetae present; gonostylus axe-shape, apical third unusually narrow, base much wider, subrectangular; hypoproct with basal process broadly triangular ( Colombia)........... O. dolabriformis Scarbrough, 2002 .

19. Female T9+10 and genital fork fused as a wide, dark sclerotized bridge ventrally; T9+10 folded laterally forming a prominent keel; genital fork with 2 large, oval depressions present medially; S8 with apical margin strongly and abruptly produced medially; spermatheca oval ( Bolivia)............................................... O. tropidus Scarbrough, 2002 .

19’. Female T9+10 and genital fork normal; T9+10 apical corner pointed, genital fork membranous, only small depressions present; S8 with apical margin only narrowly produced medially; spermatheca wider basally than apically or spherical........... 20

20. Male terminalia wider than long, epandrium with a wide notch along dorsoapical margin, and angled obliquely posteriorly; gonocoxal macrosetae unusually long and pointed; gonostylus with strong right angled basal process; phallus apically wide, angled ventrally, parameral sheath hood-like; female S8 with apical margin medially broadly produced, sides oblique; spermatheca spherical ( Peru)................................................. O. tanpadiensis Scarbrough, 2002 .

20’. Male terminalia longer than wide, epandrium only slightly notched along dorsoapical margin, and horizontal, not especially angled posteriorly; gonocoxal macrosetae shorter, apex round; gonostylus flat, without prominent lateral process basally; phallus horizontal, parameral sheath flared preapically, not hood-like; female S8 with apical margin slightly produced medially, narrow with almost vertical sides; spermatheca wider basally than apically ( Venezuela)......... O. flexus Scarbrough, 2002 .

21. Hind femur with most or all ventral macrosetae brown....................................................... 22

21’. Hind femur with most or all ventral macrosetae yellow...................................................... 24

22. Hind coxa entirely, fore and mid coxae apically usually red to brown yellow; femora yellow-brown with reddish tint, darkest apically, lighter yellow to reddish yellow basally and posteriorly; hind femur basally with 1–2 contrastingly stout posteroventral macrosetae; epandrium abruptly narrow apically; gonocoxite with sparse setae; spermatheca elliptical ( Guatemala to Panama).......................................................................... O. angustatus Scarbrough, 2002 .

22’. Coxae darker, brown to black; femora never reddish, dull yellow-brown anteriorly and dorsally or mostly yellow; hind femur basally without contrastingly stout posteroventral macrosetae; epandrium not abruptly narrowing apically; gonocoxite with an unusually thick macroseta; spermatheca spherical or ovate.................................................... 23

23. Hind femur dull yellow-brown dorsally and anteriorly, yellow elsewhere, most anteroventral macrosetae yellowish; mid femur of male with preapical, posterodorsal macrosetae absent, 5–6 long, posteroventral macrosetae present, none beyond middle; epandrium wide apically, pointed; gonocoxite with two digitate processes on inner corner, one hooked with a moderate thick, acuminate, black macroseta; female S8 rectangular apically, spermatheca spherical ( Peru).... O. destitutus Scarbrough, 2002 .

23’. Hind femur mostly yellow, brown-yellow to brown apically, darkest dorsally, anteroventral macrosetae brown; mid femur with preapical, posterodorsal macrosetae, male with only long, posteroventral setae basally, short, posteroventral macrosetae often present on apical third; epandrium triangular apically, gradually tapering to a pointed apex; gonocoxite with inner corner wide, margin oblique with one short, thick macroseta; female S8 with medioapical margin curved anteriorly, sides oblique; spermatheca ovate ( Venezuela)................................................ O. triangularis Scarbrough, 2002 .

24. Femora dorsally and at least dorsal half anteriorly dark brown, ventrally and posteriorly yellow, demarcation between colors abrupt............................................................................................. 25

24’. Femora with colors more diffuse, brown to yellow transition gradual over a wider area, never abrupt ( Venezuela).................................................................................... O. fernandezi Scarbrough, 2002 .

25. Epandrium wide apically, apex pointed, gonocoxite with a large, flat, “fang-like” macroseta; female with medioapical margin of S8 curved forward, sides oblique ( Venezuela).................................. O. unguiculatus Scarbrough, 2002 .

25’. Epandrium narrow apically, apex round, gonocoxite with sparse setae only; female with medioapical margin of S8 rectangular, sides almost parallel ( Bolivia, Argentina)....................................... O. tucumanensis Scarbrough, 2002 .

Kingdom

Animalia

Phylum

Arthropoda

Class

Insecta

Order

Diptera

Family

Asilidae

SubFamily

Ommatiinae

Genus

Ommatius

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