Notagonum reidi, Baehr & Reid, 2017

Baehr, Martin & Reid, Chris A. M., 2017, On a Collection of Carabidae from Timor Leste, with Descriptions of Nine New Species (Insecta: Coleoptera, Carabidae), Records of the Australian Museum 69 (6), pp. 421-450 : 432-434

Notagonum reidi sp. nov. Baehr

Figs 5 , 14, 21

Holotype ♂, “ TIMOR LESTE Mt Laritame 8.69178S 126.38719E water tank, edge moss forest, 1150m TL2012/ 011/016 b/light/ bucket trap 31.v.2012 ” K402620 ( AMS) . Paratypes (22): 4 ♂♂ and 7 ♀♀ same data as holotYpe: K402603, K402609–611, K402613–614, K402617–619 ( AMS) ; 1 ♂, 1 ♀ K402605, K402612 ( CBM) ; 4 ♂♂ and 1 ♀, “ TIMOR LESTE Mt Laritame 8.69181S 126.38939E water tank, edge moss forest, 1250m TL2012/012/036 black light/ bucket trap 31.v.2012 ” K402621–625 ( AMS) ; 1 ♀, “ TIMOR LESTE Mt Laritame 8.69178S 126.38719E water tank, edge moss forest, 1150m TL2012/011/030 yellow pans 30.v.2012 ” K402606 ( AMS) ; 1 ♀, “ TIMOR LESTE Mt Laritame 8.69181S 126.38939E water tank, edge moss forest, 1250m TL2012/012/023 yellow pans 31.v.2012 ” K402607 ( AMS) ; 1 ♀, “ TIMOR LESTE Hatubuilico Rd 11.5km W t’off h’waY 8°53'12"S 125°32'49"E E urophYlla /Vacc pasture/ gully 2050m 24–26.v.2012 J.Recsei TL2012/080/? Pitfall ” K402626 ( AMS) ; 1 ♂, “ TIMOR LESTE Hatoudo 8°58'56"S 125°37'27"E riverine rainforest, 735m 26.v.2012 C. Reid TL2012/088/546 mv light sheet” K402615 ( AMS) ; 1 ♂, “ TIMOR LESTE Bobonaro 9:01'47.6"S 125:19'29.1"E at light, hotel balcony 835m 28.xi.2011 V. Kessner Z. Afranio TL169/11” K402604 ( AMS) ; 1 ♂, “ TIMOR LESTE Mt Laritame water tank 5; - 8.69178S 126.38719E; 31 May 2012; 1150m;edge of moss forest; D.J.Bickel, A.Mitchell, J.B.Weiner, V.Kessner, A. V. Ribeiro, TL2012/011/016 [black light bucket trap]” K402602 ( AMS) .

Etymology. The name, proposed by MB, acknowledges co-author and collaborator Dr Chris Reid, who prepared this material.

Diagnosis. A medium sized, more or less dark piceous species without distinct spines or denticles at the apex of the elytra; distinguished from N. pleurale ( Jordan, 1894) by slightly lesser body size, indistinct dark red spots on the frons, impunctate thoracic and abdominal sterna, and almost similarly sized lobes of the 4th tarsomere of the metatarsus; and from N. angusticolle sp. nov. by wider pronotum with coarsely punctate lateral part of the base, shorter elytra, shorter antenna, and different structure of the internal sac of the aedeagus.

Description. Measurements. Body length: 7.2–8.4 mm; width: 2.85–3.3 mm; ratios: width/length of pronotum: 1.29– 1.32; width widest diameter/base of pronotum: 1.19–1.22; width base/apex of pronotum: 1.23–1.25; width pronotum/ head: 1.30–1.35; length/width of elytra: 1.62–1.65; length/ width of 6th antennomere: 2.55–2.65.

Colour ( Fig. 5 ). Dark piceous to almost black, lateral margin of the pronotum inconspicuously, but rather widely, more or less pale red, margin of elytra inconspicuously, extremely narrowly dark red. Labrum and mandibles reddish-piceous, palpi and antenna more or less pale red, 1st antennomere at least at the undersurface paler than the rest. Frons with two large though indistinct, more or less dark red spots. Legs dark red to reddish-brown, outer surface of the tibiae slightly darker. Epipleura of prothorax and elytra more or less pale red, lower surface reddish-brown.

Head ( Fig. 5 ). Of average size, dorsal surface moderately convex. Eye large though laterad but moderately protruded; orbit short, oblique, about one third of length of eye. Labrum anteriorly straight; mandibles of normal size and shape; palpi elongate, impilose. Mentum with acute, unidentate tooth and two elongate setae. Submentum quadrisetose. Antenna elongate, median antennomeres slightly> 2.5 × as long as wide, densely pilose from middle of 4th antennomere. Frons slightly convex, frontal furrows linear, rather shallow, slightly turned outwards. Usually mediadlly of eye with a more or less distinct, shallow, circular groove. Neck with shallow but distinct transverse sulcus. Surface glabrous, without microreticulation except on labrum, with rather sparse, extremelY fine punctures that are onlY visible at verY high magnification, verY glossY. OnlY laterad of the frontal furrows surface slightly rugose.

Pronotum ( Fig. 5 ). Rather wide, widest at or slightly in front of middle, dorsal surface gently convex. Base rather wide, wider than apex. Apex well excised; apical angle protruded and widely rounded at tip; lateral margin in anterior half gently convex, in basal half oblique and straight or but slightly convex, near base faintly sinuate. Base in middle straight, laterally slightly oblique. Basal angle obtusely angulate, c. 100°. Apex and middle of base margined, lateral parts of base barely margined. Lateral margin and channel rather wide and deep, basad widened and explanate. The anterior transverse sulcus fairly deep, narrow, the posterior sulcus wide and shallow; median line distinct, well impressed, almost complete. The anterior marginal seta situated slightly in front of middle, the posterior seta situated at basal angle. Disk with irregularly transverse, superficial strioles. The lateral margin in basal half and the lateral parts of the base with coarse, rugose punctures. Disk with extremelY fine and sparse punctures barelY recognizable even at high magnification, and with verY superficial traces of transverse microreticulation, surface very glossy.

Elytra ( Fig. 5 ). Moderately short and wide, slightly widened apicad, widest slightly behind middle, dorsal surface moderately depressed. Humerus wide, slightly produced but widely rounded, lateral margin in basal half almost straight but slightly oblique, in apical half regularly convex, with rather shallow, not angulate excision near apex, margin slightly incurved towards suture and without, or with a faint, obtuse protuberance at suture. Lateral margin very narrow. Striation complete and well impressed, all striae extended to apex, impunctate. Intervals depressed. The anterior discal puncture situated at basal fourth and attached to the 3rd stria, the median puncture situated slightly behind middle, the third puncture about at apical fifth, both, median and posterior punctures attached to the 2nd stria. Setae fairly elongate. Marginal series consists of 21–22 punctures that are not or barelY interrupted in middle. Surface with extremelY fine, barely recognizable punctures, without microreticulation or with finest traces of transverse microreticulation, verY glossy, slightly iridescent.

Lower surface. Surface, including abdomen, impilose and almost impunctate, except the narrow mesepisternum which is punctate. Surface glossy throughout. Metepisternum elongate, c. twice as long as wide at apex . Terminal sternum in male bisetose, in female quadrisetose.

Legs. Of normal shape and structure. Tarsi sulcate on both sides. 4th tarsomere of metatarsus little excised, the outer lobe little longer than the inner one. 1st–3rd tarsomeres of the male protarsus widened and biseriately squamose.

Male genitalia ( Fig. 14). Genital ring rather narrow, laterally slightly convex, with moderately wide asymmetric, obtusely convex apex and very narrow base.Aedeagus rather narrow and elongate, straight, depressed, lower surface, apart from near base, almost straight.Apex rather elongate, straight, depressed, regularly triangular. Internal sac with several sinuate, sclerotized rods. Both parameres comparatively short and compact, the right one stouter, with obliquely rounded apex, the left one with obtusely triangular apex.

Female gonocoxites ( Fig. 21 ). Gonocoxite 1 large, with about 8 stout ensiform setae at the apical rim of the ventral surface; gonocoxite 2 comparatively small, slightly curved, with obtusely angulate apex, with one elongate dorsomedian ensiform seta above middle, two or three rather stout ventro-lateral ensiform setae in middle, and two short, preapical nematiform setae originating from a pit. Lateral plate along the whole apical margin with many short and rather delicate setae.

Variation. Very little variation noted,

Distribution. Upland areas of Timor Leste, from Bobonaro to Mount Laritame.

Collecting circumstances. Most specimens were collected in or near rain forest and moss forest between 735 and 1250 m, usually at light or in yellow pans, one, however, in an Eucalyptus urophyllum-Vaccinium pasture gully at 2050 m.

Figures

Figure 5. Notagonum reidi sp. nov. (body length 7.2 mm).

Figures 18–22. Female gonocoxites 1 and 2. (18) Mecyclothorax timorensis sp. nov. (scale 0.1 mm). (19) Rhytisternus externus sp. nov. (scale 0.5 mm). (20) Arhytinus timorensis sp. nov. (scale 0.25 mm). (21) Notagonum reidi sp. nov. (scale 0.25 mm). (22) Aristolebia timorensis sp. nov. (scale 0.5 mm).