Penthostola subnigrantis, Groenen & Wangdi, 2019

Groenen, Frans & Wangdi, Karma, 2019, The Tortricidae (Lepidoptera) of Bhutan, Zootaxa 4658 (3), pp. 526-540 : 532

Penthostola subnigrantis , sp. nov.

Figs. 22 , 49

Holotype. ♂, Bhutan, Chhukha, Gedu , 26º55´14´´N, 89º31´17´´E, 2030 m, 5.viii.2016, sta. 8, C. Gielis, genitalia slide FG3334 ( NBCS).

Diagnosis. Penthostola subnigrantis is similar to P. nigrantis Kawabe, 1995 in colour and male genitalia. It differs from P. nigrantis in the rounded top of the gnathos, the group of spines at the costal prominence, and the longer spines at the ventral prominence of the valva.

Description. Male. Head ( Fig. 22 ): Vertex and scape of antenna grey-brown, labial palps ca. 2 times diameter of compound eye, second segment basally grey-brown, apically brown, third segment grey-brown. Thorax: Nota and tegulae grey-brown; forewing length 7 mm; forewing subrectangular, costa curved, termen oblique, dorsum straight. Ground colour brown, overlaid with black, a semi-ovoid costal spot from 2/5 to just before apex, mixed black and grey, at the basal margin of this spot a white dot, two blue-grey spots at middle of dorsum. Fringe concolourous with wing. Hindwings shiny brown-fuscous, darker apically. Fringe grey with a darker subbasal band. Abdomen: Genitalia ( Fig. 49 ) with tegumen tall, uncus rudimentary, socii pendant and long haired, gnathos short with rounded top; valva slender, excavation large, a few short setae on innerside, sacculus simple with long setae, cucullus with basal, costal, and ventral prominences; at ventral prominence three long spines, along inner margin of this prominence a group of dense setae; costal prominence smaller with a group of ca. one-half length of three ventral spines; lobe of cucullus setose; phallus slender, slightly tapering, ca. as long as valve. Female unknown.

Distribution. Bhutan.

Etymology. The name subnigrantis refers to the close relationship with P. nigrantis .

Figures

FIGURES 19‒36. Aduts and male genitalia of Cochylini, Archipini, Bactrini, Olethreutini, Eucosmini. 16. Meridemis bathymorpha, male. 17. Meridemis bathymorpha, female. 18. Borneogena trashiyana, female holotype. 19. Isodemis illiberalis, male. 20. Bactra cophinana, female holotype. 21. Lobesia ambigua, male. 22. Penthostola subnigrantis, male, holotype), 23. Metendothenia ordospina, male. 24. M. brunnofasciana, female holotype. 25. Peridaedala nigrifasciana, female holotype. 26. Gibberifera glaciata, male. 27. Crocidosema plebejana, female. 28. Lepteucosma charassuncus, male. 29. Epiblema charadrias, male. 30 E. charadrias, female. 31. E. albulusana, male holotype. 32. Microsarotis bicincta, female. 33. Eupoecilia jakarana, male genitalia, holotype. 34. Eupoecilia jakarana, phallus. 35. E. gedui, male, holotype. 36. E. gedui, phallus.

FIGURES 37‒55. Male and female genitalia of Tortricini, Archipini, Olethreutini, and Eucosmini. 37. Archips limatus, male. 38. A. limatus, phallus. 39. Chirapsina expleta, male. 40. C. expleta, phallus. 41. Clepsis humana, male. 42. Lumaria phuntschona, male. 43. L. phuntschona, phallus. 44. Meridemis bathymorpha, male. 45. M. bathymorpha, phallus. 46. Isodemis illiberalis, male. 47. I illiberalis, phallus. 48. Lobesia ambigua, male. 49. Penthostola subnigrantis, male holotype. 50. Metendothenia ordospina, male. 51. Gibberifera glaciata, male. 52. Lepteucosma charassuncus, male. 53. Epiblema charasdrias, male. 54. Epiblema albulusana, male holotype. 55. Acleris perfundana, female.