Misgolas fredcoylei, Wishart & Rowell, 2008

Wishart, Graham & Rowell, David M., 2008, Trapdoor Spiders of the Genus Misgolas (Mygalomorphae: Idiopidae) from Eastern New South Wales, With Notes on Genetic Variation, Records of the Australian Museum 60 (1), pp. 45-86 : 71-72

Misgolas fredcoylei n.sp.

Figs 16A–F , 24D

Type material. HOLOTYPE?, AM KS36704 , Chichester State Forest ( RM), Berrico Road, approximately 2 km north of intersection with Kunungra Road (32°05'S 151°45'E), 1,030 m, 4 Feb.–9 Apr. 1993, pit fall trap site 35BR, M. Gray, G. Cassis . PARATYPES ??, AM KS38604 , Chichester State Forest ( RM), Berrico Road, 100 m north of trig. tower (32°06'22"S 151°45'51"E), 940 m, 4 Feb.–9 Apr. 1993, pit fall trap site 35BM, M. Gray, G. Cassis . AM KS86215 & AM KS86219 , data as for holotype .

Other material examined. Adult! and juvenile, AM KS38605 , data as for paratype AM KS38604 .

Diagnosis. In male: medium sized brown spiders, carapace length c. 6.5–7.5; retrodorsal surface of metatarsi IV with spines, at least one midway ( Fig. 16F ); venter entirely pallid ( Fig. 16E ). Palpal bulb ( Fig. 16B,C ) with retrolateral embolic flange with c. 12 fine folds; embolus short, apophysis absent. Conformation of palp as figured ( Fig. 16A,D ); cymbium with long blunt stick-like spines inclined c. 45° forward, retrolateral tibial apophysis not basally bent downwards, distal tibial apophysis small; large black subdistal spine projecting from proventral surface of tibial excavation ( Fig. 16D ). Female not known.

Description

Male holotype ( Fig. 16A–F ). Size. Carapace length 6.51, width 5.34. Abdomen length 5.46, width 3.43. Colour. In alcohol carapace, legs, palps & chelicerae light tan; dark brown smudges absent from lateral limb surfaces. Few sparse pallid hairs on caput, absent on limbs.Abdomen dorsum light brown mottled with dark brown; “chevrons” absent. Venter entirely pallid. Carapace. Edge fringed with black bristles some extending onto post foveal surface; some smaller bristles on remaining caput area. Line of 6 bristles (some rubbed off) on caput arch. Group of three bristles between PME; group of 9 on clypeus. Area extending onto pleuron membrane below clypeus narrow without setae. Fovea width 0.99, straight, posterior wall not pitted. Eyes. Raised on mound. Areas between adjacent eyes black. Widths of anterior and posterior rows c. equal, 1.08 and 1.07 respectively. Eye group length 0.61, width/length ratio 1.75. Line joining posterior edge of ALE transects anterior fifth of AME. Posterior row straight in front, recurved behind. Chelicerae. Rastellum first row 6(6) strong spines; c. 11(8) behind and extending along pd cheliceral edge. Intercheliceral tumescence elongate, faint, not distinct. Fang groove with 10(7) promarginal teeth and 9(10) smaller retromarginal/intermediate row teeth. Labium. Bulbous, length 0.61, width 1.13. Labio-sternal suture undivided, broad narrowing laterally. Maxillae c. 8(13) stick-like anteroental cuspules. Sternum. Length 3.25, width 2.45. Two pairs small round sigilla separated from margin by their own diameter; posterior third pair absent (present in paratypes). Legs. Tibia I with distal bifid apophysis; distal process with 2(2) short blunt spines, proximal process with 3(3) short pointed spines.

palp I II III IV femur 3.50 6.38 6.01 5.09 6.69 patella 1.66 3.01 2.89 2.39 3.00 tibia 3.44 4.67 4.61 3.25 5.65 metatarsus — 4.55 4.30 4.17 5.77 tarsus 1.60 2.82 2.70 2.64 3.19 total 10.20 21.43 20.51 17.54 24.30

Palp ( Fig. 16A ). Cymbium with c. 38 long, blunt, stick-like spines, each terminated by a posteriorly positioned point, anteriorly inclined c. 45° to cymbium axis and distributed over distal third of d surface and distal two-thirds of pd surface. RTA broad without swellings, d and rd surface covered with c. 45 squat pointed spines; proximal half of rv edge of tibial excavation with c. 30 similar spines; distal half of rv edge with c. 16 longer spines extending onto small straight DTA. TEM prominent, pallid, adjacent to RTA. TET not seen (discernible on paratype AM KS38604). Large black subdistal spine (fractured on right palp) projecting from pv surface of tibial excavation ( Fig. 16D ). Bulb. ( Fig. 16B,C ) Embolic rl flange with c. 10 fine folds, margin convexly curved, folding under short embolus; embolic apophysis absent. Scopula. Dense and complete on tarsi I and II, weak and incomplete on metatarsi I and II, absent on legs III and IV. Trichobothria. Palp: tarsi 7, tibia pd 5, rd 6. Leg I: tarsi 10, metatarsi 11, tibia pd 6, rd 5. Leg II: tarsi 10, metatarsi 10, tibia pd 6, rd 7. Leg III: tarsi 10, metatarsi 11, tibia pd 6, rd 6. Leg IV: tarsi 11, metatarsi 13, tibia pd 6, rd 6. Leg spination. Leg I: tarsi v 1 subdistal, rv line of 4 adjacent scopula; metatarsi pv 011, rv 0112; tibia v 0112. Leg II: tarsi pv 2 subdistal, rv line of 6 adjacent scopula; metatarsi pv 00111, rv 02132; tibia v 01112, pd 0110. Leg III: tarsi v 10 scattered, rl 01110; metatarsi v 15 scattered, d 002220; tibia v 00112, pl 0110, rl 0110; patella pd 7. Leg IV: ( Fig. 16F ) tarsi v16 scattered; metatarsi v 10 scattered, rd 010; tibia v 00113. Abdomen. Dorsum with cover of long brown bristles. Hairs and short soft bristles on lateral surfaces. Longer weak bristles on venter.

Remarks. The female and juvenile specimens AM KS38605 referred to as “Other material examined” were collected from the same pitfall trap as paratype AM KS38604. Although these specimens are possibly the same species as the holotype they are not recognized as M. fredcoylei in this work as spination of metatarsi IV does not conform with that of the male types.

Etymology. The species is named in recognition of eminent arachnologist and teacher Frederick A. Coyle, Professor of Biology, Western Carolina University, USA.

Distribution and natural history ( Fig. 24D ). This spider is known only from the Chichester State Forest approximately 25 km South West of the town of Gloucester. The burrow is unknown.

Figures

Fig. 16. Misgolas fredcoylei n.sp. A–C,E,F?, holotypeAM KS36704:(A), right palp retrolateral.(B,C), right bulb: B, dorsal; C, prolateral. D?, paratype AM KS38604: (D), right palp prolateral. (E), venter. (F), right tarsus/metatarsus IV retrolateral.

Fig. 24. A–D. Distribution of Misgolas species in the New South Wales Central Coast (eastern Australia) based on material examined (Inset [Fig. 24A] with catchment area).