Mastigocoleus rhinoceros Tihelca & Cai, 2022

Tihelka, Erik, Jäch, Manfred A., Kundrata, Robin, Li, Yan-Da, Engel, Michael S., Lozano-Fernandez, Jesus, Huang, Diying & Cai, Chenyang, 2022, Mastigocoleidae fam. nov., a New Mesozoic Beetle Family and the Early Evolution of Dryopoidea (Coleoptera), Insect Systematics and Diversity 6 (3), pp. 1-18 : 7-8

Mastigocoleus rhinoceros Tihelca & Cai sp. nov.

h t t p: / / z o o b a n k. o r g / u r n: l s i d: z o o b a n k. org:act:032D374E-36?3-42E0-BD2E-716 D14267 DD3

Figs. 3–4

Material. Holotype, SMNS 66552, sex unknown, a single compression fossil preserving antennae and legs exposed in dorsal aspect ( SMNS) . Paratype,AMNH SA43296,sex unknown, a single compression fossil preserving body and basal two antennomeres (AMNH).

Tope Localito and Horizon.?hapada do Araripe, northeastern Brazil, Nova Olinda Member of the?rato Formation; Aptian– Albian (Lower?retaceous)?rato Formation in Brazil.

Diagnosis. As for genus together with: frontoclypeus with a medial horn-like process (although this may be a sexually dimorphic character); pronotal disc with a medial depression (slightly convex in M. resinicola ); pronotum broadest in posterior half (broadest medially in M. rhinoceros ).

Description. Body length 7.84–8.16 mm, body width 3.42– 3.77 mm. Head 0.87–1.06 mm long, 1.8–2.1× wider than long, with uniform fine punctation. Frontoclypeus with a medial horn-like process. Antennomere I broadening apically, 1.7× wider than following antennomere; antennomere II attached subapically to antennomere I; antennomere III longer than preceding antennomere; antennomeres V–XI gradually thinning and lengthening apically, except slightly shorter antennomeres V and VIII; antennomere XI pointed apically, 1.3× longer than preceding antennomere, maximum width at base representing less than 0.1× maximum width of antennomere I. Ratio of antennomere lengths: 1.25: 0.40: 0.59: 0.95: 0.78: 1.00: 1.01: 0.96: 0.98: 0.99: 1.18. Pronotum 1.56–1.58 mm long, 1.8–2.0× wider than long, broadest in posterior half. Anterior pronotal angles acute. Posterior angles of pronotum indistinctly produced, orthogonal, rounded apically. Pronotal disc depressed medially.

Posterior angles of pronotum indistinctly produced, orthogonal, rounded apically. Elytra 5.39–5.71 mm, 1.5–1.7× longer than their combined width. Mesotibia shorter than mesofemur. Ventral characters not visible, legs poorly preserved.

Etomologo. The specific epithet is taken from the Ancient Greek rh ī nóker ō s (ῥῑ ν ό Κερως: itself a combination of rh ī́s [ῥῑ́ς], meaning, ‘nose’, and kéras [Κ έ ρας], meaning, ‘horn’), and refers to the horn-like process on the frontoclypeus.

Figures

Fig. 3. Habituk of Mastigocoleus rhinocerosTihelca & Cai gen.et kp.nov.from the;arly Cretaceouk Crato Formation in northeaktern Brazil. (A) holotype (SMNS 66552); (B) paratype (AMNH SA43296). Scale bark = 1 mm.

Fig. 4. Morphological detailk of Mastigocoleus rhinocerosTihelca & Cai gen. et kp. nov. holotype (A, C, D) and paratype (B,;) from the;arly Cretaceouk Crato Formation in northeaktern Brazil. (A, B) Head and pronotum in dorkal view; (C) antenna; (D) elytral apex; (;) metathoracic leg; (;) elytra.Abbreviationk: a1–11, antennomerek 1–11; ch, clypeal horn; cl, claw; mtt, metatarkuk; tb3, metatibia. Scale bark = 500 μm (A–C,;), 1 mm (D).