Aegognathus Leuthner, 1883

Ríos-Málaver, Juan Sebastián Dueñas Cáceres ı Cristóbal & Grossi, Paschoal Coelho, 2019, Taxonomic contribution on the Andean species of Aegognathus (Coleoptera: Lucanidae) with two new species from Colombia and Peru, JOURNAL OF NATURAL HISTORY 53 (35), pp. 2145-2164 : 2154-2155

Key to Andean Region Aegognathus Leuthner species

1. Body apparently uniformly blackı at least along head and pronotum but showing some small dark-red spots ventrally and dorsally with bluish or reddish reflections along elytra. ........................................................................................................................................... 2. Body with distinctly black and red to orange spots. ............................................................. 4.

2. Last Ventrite with two conical tubercles at both sidesı and dense yellowish setae. Male aedeagi slenderı strongly sclerotisedı with setose median lobe at sidesı X shaped dorsal cross bar presenting two acuminate projections near the median axis. Female unknown. ............................................... ............................................... A. waterhousei Leuthner. Last ventrite with no lateral tuberclesı presenting or not a somewhat continuous lateral carina along abdominal ventrites. Different genitalia. ............................................ 3.

3. Elytra showing bluish reflections and reddish longitudinal costae. Mandibles with a dorsalı basal carinaı inner margin wavy. Median tooth almost trianguları apex emarginateı slightly bifurcated. Aedeagus shortı weakly sclerotisedı with a somewhat flatı short everted sac. Female unknown. ................... ................... A. dulima . Body mainly black. Mandibles not wavy internallyı and with no dorsal carina at base. Females entirely blackı sometimes showing bluish reflections along elytra; elytra entirely punctateı with no traces of carina; left mandible showing a transverse laminar tooth presenting one bigger pit and two smaller at base. Abdominal ventrites presenting a lateral longitudinal carinaı borderedı resembling a continuous line along lateral margins. Male genitalia elongateı somewhat slenderı strongly sclerotisedı presenting a cylindrical and somewhat long internal sac. .................. .................. A. arnaudi .

4. Heads as wide as elytraı mandibles somewhat cylindricalı strongly curved inwards with a media tooth pointing upwards in major males (not present or reduced in small specimens)ı females with laminar median tooth; Ocular canthus angulatedı especially in femalesı sometimes rounded sometimes emarginate. ................................................... 5. Males with head and pronotum robustı wider than elytra. Mandibles shortı wide and somewhat flattenedı rectangular in cross sectionı all teeth pointing to the median axis of the bodyı never longer than mandible width; females with ocular canthi distinctly rounded. Gena strongly carinate in males and strongly punctate in females. Male genitalia with an internal sac subconical at apexı medially concave. ................................... ............................................................................................................ A. aguirei Arnaud & Bomans.

5. Body entirely bicolouredı with black and orange marks. Elytra presenting 2 – 4 orange longitudinal striaeı some costae. Male genitalia with parameres apically roundedı dorsally with concave posterior margin. ............................ A. soulai Arnaud & Bomans. Body bicoloured with black and dark red spots. Elytra without distinct costaeı even with longitudinal red striae. Genitalia different from above .............................................. 6.

6. Male mandibles with truncate apexı dorsal angle slightly elongateı rounded; preapex in medium and major males with a small tooth. Female mandibles dorsally carinate along the external and internal margins; ocular canthi quadrate. Male second abdominal ventrite without transverse wrinkles along lateral margins. ............................................. .............................................................................................................. A. similis Arnaud & Bomans. Mandibles with rounded or acuminate apexı secondı or second and third abdominal ventrites presenting wrinkles along lateral margins. ............................................................. 7.

7. Second and third abdominal ventrites presenting transverse wrinkles. Elytra uniformly punctateı punctures elongate to coalescent; colour pattern almost uniform from darkish red to black with no distinct area of costate or subcostate striae. ......................... ........................................................................................................ A. confusus Arnaud & Bomans. Only second abdominal ventrite with transverse wrinkles. Elytra bicolouredı with punctures not uniformly distributedı sometimes with longitudinal costae; elytral punctures circular and coalescent. ................................................................................................ 8.

8. Male mandibles more abruptly incurvedı with a somewhat sharp apexı with a preapical transverse longitudinal carina. Pronotum and elytra showing distinctly long yellowish setaeı densely distributed along elytra; setae longer than twice a puncture diameter; setae in females distinctly broader. Female elytra with disc bicoloured. Male genitalia with X shaped; dorsal cross bar poorly lobedı but broader at posterior portion. .............................................................. A. damasoi Arnaud & Bomans. Male mandibles gradually incurvedı with truncate to rounded apexı preapical carina absent. Yellowish setae mainly along elytraı but shortı as long as a punctate diameter. Female elytral disc almost uniformly blackı setae slenderı hair-like. Male genitalia with X-shaped dorsal crossbar with broader posterior lobes. ...... A. leuthneri Van de Poll.