Cyclocephala melanocephala (Fabricius, 1775)

Moore, Matthew R., Cave, Ronald D. & Branham, Marc A., 2018, Annotated catalog and bibliography of the cyclocephaline scarab beetles (Coleoptera, Scarabaeidae, Dynastinae, Cyclocephalini), ZooKeys 745, pp. 101-378 : 190-191

Cyclocephala melanocephala (Fabricius, 1775)

Melolontha melanocephala Fabricius, 1775: 36 [original combination].

Cyclocephala melanocephala (Fabricius) [new combination by Burmeister 1847: 56-57].

Dichromina melanocephala (Fabricius) [new combination by Casey 1915: 160].

Cyclocephala melanocephala (Fabricius) [revised combination by Arrow 1937b: 8, 13].

syn. Chalepus leucophthalmus Fischer, 1823: 265 [original combination].

Dyscinetus leucophthalmus (Fischer) [new combination by Harold 1869a: 123].

Cyclocephala melanocephala (Fabricius) [synonymy by Arrow 1911: 172].

syn. Cyclocephala dimidiata Burmeister, 1847: 57 [original combination].

Dichromina dimidiata (Burmeister) [new combination by Casey 1915: 161].

Cyclocephala (Dichromina) dimidiata (Burmeister) [revised combination and new subgeneric classification by Saylor 1937: 70].

Cyclocephala dimidiata Burmeister [removal of subgeneric classification by Arrow 1937b: 8, 10].

Cyclocephala melanocephala (Fabricius) [synonymy by Endrődi 1964: 466].

syn. Cyclocephala elegans Horn, 1871: 337 [original combination].

Cyclocephala dimidiata Burmeister [synonymy by Horn 1875: 143].

Dichromina elegans (Horn) [revalidated species status and new combination by Casey 1915: 162].

Cyclocephala dimidiata Burmeister [synonymy by Saylor 1945: 382].

Cyclocephala melanocephala (Fabricius) [synonymy by Endrődi 1964: 466].

syn. Cyclocephala rubiginosa Burmeister, 1847: 59 [original combination].

Cyclocephala melanocephala rubiginosa Burmeister [new subspecific status by Chalumeau and Gruner 1977: 584].

Cyclocephala melanocephala (Fabricius) [synonymy by Ratcliffe and Cave 2015].

syn. Cyclocephala ventralis Erichson, 1847a: 97 [original combination].

Cyclocephala melanocephala (Fabricius) [synonymy by Endrődi 1964: 466].

syn. Dichromina ocularis Casey, 1915: 162 [original combination].

Cyclocephala ocularis (Casey) [new combination by Arrow 1937b: 8, 14].

Cyclocephala dimidiata Burmeister [synonymy by Saylor 1945: 382].

Types.

Type of M. melanocephala at BMNH (Banks Collection) ( Endrődi 1966). Lectotype of C. ventralis at ZMHB ( Endrődi 1966). Type of C. dimidiata is missing ( Endrődi 1966). The type of Cyclocephala leucophthalmus is unknown ( Endrődi 1966). Type of D. ocularis at USNM ( Endrődi 1966). Type of C. elegans at USNM ( Endrődi 1966). The type of C. rubiginosa is missing ( Endrődi 1966).

Distribution.

ARGENTINA: Córdoba, Salta, Santa Fe, Tucumán. BELIZE: Cayo, Orange Walk, Stann Creek, Toledo. BOLIVIA: Beni, Cochabamba, Santa Cruz. BRAZIL: Amazonas, Bahia, Ceará, Espírito Santo, Goiás, Mato Grosso, Mato Grosso do Sul, Minas Gerais, Paraná, Pernambuco, Rio de Janeiro, Rio Grande do Norte, Rio Grande do Sul, Roraima, Santa Catarina, São Paulo. COLOMBIA: Antioquia, Atlántico, Boyacá, Casanare, Cauca, Cesar, Chocó, Córdoba, Cundinamarca, Huila, La Guajira, Meta, Risaralda, Santander, Tolima, Valle del Cauca. COSTA RICA: Alajuela, Cartago, Guanacaste, Heredia, Limón, Puntarenas, San José. DOMINICA: St. David, St. Joseph, St. Patrick, St. Paul. DOMINICAN REPUBLIC: Barahona. ECUADOR: Bolívar, Cañar, Esmeraldas, Guayas, Loja, Los Ríos, Morona-Santiago, Santa Elena. EL SALVADOR: Ahuachapán, Chalatenango, La Libertad, Morazán, Santa Ana. FRENCH GUIANA: Cayenne. GRENADA: St. Andrew, St. David, St. George, St. John. GRENADINES: Bequia, Canouan, Carriacou, Union Island. GUADELOUPE: Basse-Terre, Îles des Saintes, Marie-Galante. GUATEMALA: Alta Verapaz, Baja Verapaz, Chiquimula, El Progreso, Escuintla, Guatemala, Huehuetenango, Izabal, Petén, Quetzaltenango, Quiché, Retalhuleu, San Marcos, Santa Rosa, Sololá, Suchitepéquez, Zacapa. GUYANA: Cuyuni-Mazaruni, East Berbice-Corentyne, Pomeroon-Supe naam. HONDURAS: Atlántida, Choluteca, Comayagua, Cortés, El Paraíso, Francisco Morazán, Gracias a Dios, Intibucá, Olancho, Santa Bárbara, Yoro. MARTINIQUE: Fort-de-France, La Trinité, Le Marigot, Le Marin, Saint-Pierre. MEXICO: Aguascalientes, Baja California, Baja California Sur, Chiapas, Chihuahua, Distrito Federal, Colima, Durango, Guerrero, Hidalgo, Jalisco, Michoacán, Morelos, Nayarit, Nuevo León, Oaxaca, Puebla, Querétaro, Quintana Roo, San Luis Potosí, Sinaloa, Sonora, Tabasco, Tamaulipas, Veracruz, Yucatán, Zacatecas. MONTSERRAT. NICARAGUA: Carazo, Chinandega, Granada, Jinotega, León, Matagalpa, RAA Norte, RAA Sur, Río San Juan, Rivas. PANAMA: Bocas del Toro, Chiriquí, Coclé, Colón, Darien, Former Canal Zone, Panamá, San Blas, Veraguas. PARAGUAY: Asunción, Paraguarí. PERU: Ayacucho, Callao, Cusco, Junín, La Libertad, Madre de Dios, Pasco. PUERTO RICO. SAINT BARTHÉLEMY. SURINAME: Marowjine. TRINIDAD AND TOBAGO: Trinidad. UNITED STATES: Arizona, Arkansas, California, Kansas, Louisiana, Mississippi, Nevada, New Mexico, Oklahoma, Texas, Utah. VENEZUELA: Bolívar, Capital District, Carabobo, Mérida, Zulia.

References.

Fabricius 1775, 1801, Herbst 1790, Illiger 1802a, 1805, Dejean 1833, 1836b, Sturm 1843, Erichson 1847a, Burmeister 1847, Harold 1869b, Bertkau 1873, Crotch 1873, Austin 1880, Henshaw 1885, Bates 1888, Cockerell 1897, Horn 1871, 1894, Fall 1901, Fall and Cockerell 1907, Ohaus 1910, Leng and Mutchler 1914, Casey 1915, Bodkin 1919, Leng 1920, Arrow 1900, 1911, 1937b, Moore 1937, Guimarães 1944, Saylor 1937, 1945, 1948, Sanderson 1940, Blackwelder 1939, 1944, Paulian 1947, Blackwelder and Blackwelder 1948, Mariconi 1959, Linsley 1960, Zimsen 1964, Carrillo-S. et al. 1966, Martínez 1968a, Squire 1972, Ritcher and Baker 1974, Pike et al. 1976, Chalumeau and Gruner 1977, Cartwright and Chalumeau 1978, Chalumeau 1982, 1983, Boiça et al. 1984, Endrődi 1964, 1966, 1973a, 1985a, Ungaro et al. 1985, Gottsberger 1986, Morón et al. 1985, 1988, Sutton 1988, Dechambre 1992, Rogers 1992, Murray 1993, Thomas 1993, Lobo and Morón 1993, Deloya and Morón 1994, Morón 1979, 1994, Dupuis 1996, Poole and Gentili 1996, Charlet et al. 1997, Ratcliffe and Morón 1997, Sanchez Soto 1997, Woodruff et al. 1998, Bauernfeind 2001, Navarrete-Heredia et al. 2001, DeFoliart 2002, Peck et al. 2002, Carrillo-Ruiz and Morón 2003, Joly 2003, Luna et al. 2003, Marquet and Roquet 2003, Raguso et al. 2003, Riley and Wolfe 2003, Ratcliffe 1992c, 2002a, 2003, Restrepo et al. 2003, Pardo-Locarno et al. 2005a, Neita-Moreno et al. 2006, Ponchel 2006, Pacheco-F. et al. 2008, Múñoz-Hernández et al. 2008, Útima and Vallejo 2008, Barbosa Moreira et al. 2009, Smith 2003, 2009, Yanes-Gómez and Morón 2010, Neita-Moreno 2011, Giannoulis et al. 2012, Hay et al. 2012, Krajcik 2005, 2012, Breeschoten et al. 2013, García-López et al. 2013, Lima Nogueira et al. 2013, Pardo-Locarno 2013, Yepes-Rodriguez et al. 2013, Dutrillaux et al. 2007, 2014, Alberio et al. 2015, García-Atencia and Martínez-Hernández 2015, López-García et al. 2015, Ratcliffe et al. 2013, 2015, Deloya et al. 2014a, b, 2016, Dossey et al. 2016, Gasca-Álvarez and Deloya 2016, Mitasuhashi 2016, Peck 2016, Schrader et al. 2016, Ratcliffe and Cave 2006, 2015, 2017, Romero-López and Morón 2017.