Carneoryctes Özdikmen, 2009

Allsopp, Peter G. & Hutchinson, Paul M., 2020, Niuailan, a new genus to accommodate Carneoryctes peterseni (Endrődi, 1967) (Coleoptera: Scarabaeidae: Dynastinae: Pentodontini) from New Ireland, Papua New Guinea, Zootaxa 4860 (1), pp. 135-145 : 137-139

Carneoryctes Özdikmen, 2009

Cryptoryctes Carne, 1957: 154 (preoccupied by Cryptoryctes Reed, 1954 , Mammalia).

Carneoryctes Özdikmen, 2009: 139 (replacement name).

Type species. Pseudoryctes tectus Blackburn, 1892 , by original designation.

Redescription. Male (from Carne 1957; Endrődi 1985; and P.M.H. specimens) ( Figs. 3–13 ). Body reddish brown, bicolorous, or black, 12–27 mm long. Mentum strongly compressed and reflexed at ligula; maxillary palp slender, apical palpomere truncate; galea reduced to small, untoothed pieces; labrum usually visible; mandibles visible. Antenna with 10 antennomeres, club longer than shaft. Clypeus at angle to plane of frons, nearly always verti- cal, concave and with truncate anterior face glabrous or setose, disc rarely setose. Clypeofrontal ridge transverse, strongly elevated. Frons frequently setose; ocular canthi large, rounded or subangulate, densely setose. Pronotum with anterior margin corneous and bearing median elevated process or horn, latter tapering to a simple, bifid, trifid, or branched apex; anterolateral angles acute or obtuse; lateral ridges sometimes contracted or interrupted just before basolateral angles, their margins frequently with dorsal setae; basal ridge sometimes continuous; disc often setose, transversely rugulose with deep excavation limited by elevated angles or lateral horns, latter usually simple, disc rarely depressed below level of basal margin. Each elytron with sutural stria linear or linear-punctate, disc with welldefined striae, punctures simple or weakly umbilicate, surface sometimes microreticulate, epipleuron with short lateral setae continuous to apices. Pygidium setose across base, apical ridge glabrous or setose. Postcoxal prosternal process short, setose, often not visible. Abdominal sternites setose across posterior margins. Protibiae tridentate with spur inserted anterior to base of medial denticle; protarsomere 5 usually longer than protarsomere 1, not thick- ened; prothoracic claws simple, slender; metathoracic legs slender or moderately short; metatibiae bicarinate, distal ciliae short, sharp, closely set, spurs unequal, curved; metatarsi elongate, tarsomeres 1 and 5 subequal in length. Parameres symmetrical, external margin broadening pre-apically, in lateral view face arcuate, ratio of phallobase length to paramere length 1.5–2.5:1.

Included species. General geographic ranges (all in Australia) were taken from Carne (1957, 1981), Allsopp & Lloyd (1987), and Howden & Malý (2005).

Carneoryctes ater (Lea, 1917) —southwestern South Australia

Carneoryctes brittoni ( Carne, 1957) —inland-western Western Australia

Carneoryctes griseopilosus (Lea, 1917) —southeastern South Australia

Carneoryctes minchami ( Carne, 1981) —central-eastern South Australia

Carneoryctes monstrosus (Blackburn, 1895) —central-western Western Australia

Carneoryctes nigripennis (Lea, 1917) —inland southern Queensland

Carneoryctes pimbus ( Carne, 1957) —southern Northern Territory

Carneoryctes psilus ( Carne, 1957) —inland and central-western coastal Western Australia

Carneoryctes semicalvus (Lea, 1917) —southern South Australia

Carneoryctes sulcatus (Arrow, 1914) —southern Northern Territory

Carneoryctes tectus (Blackburn, 1892) —western Victoria and New South Wales, southern South Australia and Northern Territory, and inland northwestern Western Australia

Carneoryctes tricornutus ( Howden & Malý, 2005) —southern Northern Territory

Carneoryctes trifidus (Blackburn, 1895) —inland southern Queensland, northern New South Wales, and northwest- ern Western Australia

Carneoryctes truncatus ( Carne, 1957) —inland Queensland

Carneoryctes wingarus ( Carne, 1957) — Western Australia (unknown type locality) and southwestern Northern Ter- ritory

Figures

FIGURES 3–7. Carneoryctes tectus (Blackburn, 1892), male in PMH: 3, dorsal view; 4, ventral view; 5, lateral view; 6, para- meres frontal view; 7, parameres and phallobase lateral view.

FIGURES 8–13. Carneoryctes males in PMH: 8, C. brittoni (Carne, 1957), lateral view; 9, C. brittoni, dorsal view; 10, C. psilus (Carne, 1957), lateral view; 11, C. psilus, dorsal view; 12, C. wingarus (Carne, 1957), lateral view; 13, C. wingarus, dorsal view.