Dichaetophora javaensis Yang & Gao, 2017

Yang, Jin-Hua, Toda, Masanori J., Suwito, Awit, Hashim, Rosli & Gao, Jian-Jun, 2017, A new species group in the genus Dichaetophora, with descriptions of six new species from the Oriental region (Diptera, Drosophilidae), ZooKeys 665, pp. 121-146 : 130

Dichaetophora javaensis Yang & Gao sp. n.

Figs 3N-P , 4U-Y , 9

Type material.

Holotype. ♂ (#03892), INDONESIA: Cikaniki, Mt. Halimun, West Java, 7.x.2009, MJ Toda (MZB).

Paratypes: same as holotype except for 6.xi.2009 (1♀: #03887, MZB; 2♀: #03888-9, KIZ); same as holotype except for 10.xi.2009 (1♀, SEHU); Cibodas, West Java, Indonesia, 16.xi.2013, MJ Toda (1♂, SEHU); Mt. Patuha, Sugihmukti, West Java, Indonesia, 13.x.2004 (1♂, SEHU).

Diagnosis.

Postocellar setae present; spermathecal capsule somewhat dome-shaped, apically roundish; introvert 2/5 as deep as capsule height (Fig. 9J ); hypandrium sparsely pubescent in small, medial patch on caudolateral plate (Fig. 9G ); COI = C and T at sites 142 and 205, respectively; COII = T, C, C, C, G and C at sites 18, 69, 270, 303, 381 and 393, respectively (Table 5 ).

Description.

Head (Figs 3N, O , 4U-Y ): First flagellomere grayish brown. Dorsolateral tentorial apodemes slightly divergent in basal 2/5 but strongly divergent in distal 3/5; supercervical setae 18-22 per side; postocular setae 19 per side. Palpus with one subprominent, lateromedial seta. Cibarium with ca. 90 medial and 10-12 posterior sensilla per side.

Wings (Fig. 3P ) slightly fuscous; veins yellowish brown to brown. Halter and stem gray.

Legs (Fig. 3N ): All femora blackish brown; foreleg coxa, tibia and tarsus (except 5th tarsomere) grayish brown; rest grayish yellow. Apical setae present on foreleg and mid-leg tibiae. Mid-leg first tarsomere as long as total length of four succeeding tarsomeres; hindleg first tarsomere longer than total length of four succeeding tarsomeres.

Male terminalia (Fig. 9A-G ): Epandrium pubescent except for anterior margin, with 2-3 dorsal and 9-10 ventral, long setae per side. Surstylus with 2-3 short spines and medial patch of pubescence on outer surface and 10-11 recurved spines on inner, caudal portion. Tenth sternite lingulate, slightly curved. Cercus pubescent except for dorsal to caudal margin, with 14-17 long setae; sclerotized, caudoventral bridge of cerci medially with narrowly elongated process. Paramere somewhat triangular in lateral view, apically round, with sensilla on inner surface. Aedeagus apically trilobed, densely hirsute on outer lobes, dorsosubapically with a pair of small, marginally serrated flaps.

Female terminalia (Fig. 9H-J ): Oviscapt valve dorsomedially narrowly extended, with 4-5 lateral and 12-15 marginal, peg-like ovisensilla. Spermathecal capsule without spinules on outer surface.

Measurements (in mm): BL = 1.94 in holotype (range in 3♀ paratypes: 2.08-2.50), ThL = 0.86 (0.97-1.03), WL = 1.81 (1.99-2.07), WW = 0.82 (0.86-0.92).

Indices: FW/HW = 0.56 (3♀, or less if noted, paratypes: 0.49-0.52), ch/o = 0.21 (0.34-0.41), prorb = 0.76 (0.72-0.79), rcorb = 0.24 (0.29-0.31), orbito = 0.60 (0.54-0.69), dcl = 0.74 (1♀: 0.71), sctl = 0.81(1♀: 0.81), sterno = 0.60 (0.50-0.54), dcp = 0.52 (0.53-0.60), sctlp = 0.71 (0.52-0.71), C = 1.85 (1.78-1.98), 4c = 1.54 (1.35-1.40), 4v = 2.62 (2.26-2.28), 5x = 2.28 (1.73-1.91), ac = 3.03 (2.95-3.24), M = 0.76 (0.71-0.90), C3F = 0.58 (0.50-0.60).

Etymology.

Pertaining to the type locality.

Distribution.

Indonesia (West Java).

Figures

Figure 3. Left lateral habitus, head and thorax (dorsal view), wing (left, ventral view), and abdomen (dorsal view). A-D Dichaetophora tirlobita sp. n. (# 03877) E-G D. heterochroma sp. n. (# 03879) H-J D. flatosternata sp. n. (# 04172) K-M D. borneoensis sp. n. (# 03895) N-P D. javaensis sp. n. (# 03892) Q-S D. sumatraensis sp. n. (# 03890). Scale bars: 1.0 mm.

Figure 4. Head (anterior view), postocciput, palpus, and prementum (ventral and lateral view, respectively). A-E Dichaetophora tirlobita sp. n. (# 03877) F-J D. heterochroma sp. n. (# 03879) K-O D. flatosternata sp. n. (# 04172) P-T D. borneoensis sp. n. (# 03895) U-Y D. javaensis sp. n. (# 03892) Z-D D. sumatraensis sp. n. (# 03890). Scale bars: 0.1 mm except for A, F, K, P, U, Z (0.5 mm).

Figure 9. Dichaetophora javaensis sp. n. (A-G # 03892 H-J paratype # 03888). A, B Periphallic organs (posterior and posterolateral view, respectively) C surstyli and ventral portions of cerci D-F phallic organs (ventral, ventrolateral and lateral view, respectively) G distal portion of hypandrium (posteroventral view), showing a pair of small patches of sparse pubescence on the caudolateral plates (red arrows) and the paramedian setae H, I oviscapt (lateral and ventral view, respectively) J spermatheca (lateral view). Scale bars: 0.1 mm.