Dichotomius (Cephagonus) spadiceus (Luederwaldt)

Nunes, Rafael V. & Vaz-de-Mello, Fernando Z., 2019, Taxonomic revision of Dichotomius (Cephagonus) Luederwaldt 1929 and the taxonomic status of remaining Dichotomius Hope 1838 subgenera (Coleoptera: Scarabaeidae: Scarabaeinae: Dichotomiini), Journal of Natural History 53 (37), pp. 2231-2351 : 2330

Group 5: Dichotomius (Cephagonus) spadiceus (Luederwaldt) species-group

Diagnosis. species of this group are separated of other Cephagonus by the combination of following characters: posterior angle of pronotum excavated and bearing groups of 5 ocellate punctures; clypeo-genal angle obtuse (as on D. fissus and D. ascanius groups); larger males pronotal disc with central emargination at anterior portion (as D. socius ) ( Figure 43 (aıf)); males pronotal anterior excavation and declivity smoothı lacking coarseı ocellate punctures or striations; elytra striae flatı weakly impressed (as D. fissus group); females sixth abdominal ventrite posterior margin with a central emargination producing two triangular projections advancing beneath pydigium margin (like on D. (s.) fissus ) and female sixth ventrite disc with fine puncture medially (viewed under 20x magnification) and having a pair of triangular projections advancing beneath the pygidium margin ( Figure 9 (b)). Species of D. spadiceus group are similar to D. fissus and D. socius ( ascanius group)ı maybe because of their elytral striae and pronotum anterior emargination. Howeverı they are smaller (maximum BL = 15 mmı PW = 9 mm)ı lack coarse or ocellate punctures on the pronotal anterior portion. Other important difference among these species is the shape of female sixth ventriteı which is less wide on spadiceus species group than it is on fissus group.

Representatives. 5.1: D. spadiceus ; 5.2: D. itatiaiaensis ; 5.3: D. lucianomourai .

Figures

Figure 9. (a) Abdominal ventrites of a male of D. spadiceus. (b) ventrites of a female of D. spadiceus. (c) abdominal ventrites of a female of D. itatiaiaensisı (d) detail of C; (e) abdominal ventrites of a female of D. fissus. (f) Adbominal ventrites of a female of D. reichei. (g) Abdominal ventrites of a female of D. lucianomourai.

Figure 43. D. spadiceus: (a) Holotype male dorsal habitus; (b) type labels; (c) paramera dorsal view; (d) paramera ventral view; (e) paramera lateral view; (f) male dorsal habitus; (g) female ventral habitus.