Dolerus (Equidolerus) pseudoanticus Malaise, 1931

Heidemaa, Mikk, Smith, David R. & Shinohara, Akihiko, 2012, Taxonomy of Dolerus subfasciatus auct. and D. subfasciatus F. Smith with notes on the sawfly subgenus Equidolerus (Hymenoptera, Tenthredinidae), Zootaxa 3525, pp. 1-17 : 4-5

Dolerus (Equidolerus) pseudoanticus Malaise, 1931 , spec. rev.

Table 1 ; Figs 5 , 13–14 , 17

Dolerus pseudoanticus Malaise, 1931: 14 –15. Type locality: Klutchi [= Kluchi, Kamchatka, Russia]. Lectotype Ƥ here designated; right antenna missing, left half of the ovipositor (lance+lancet) glued on paper and pinned with the specimen; NHRS. Labelled: "570" [pink, printed], "KAMTSCHATKA Malaise" [pale, framed, printed], " Typus " [red, framed, printed], "[pale, framed] Dolerus pseudoanticus n.sp. (Typus)" [handwritten] Malaise det. [printed]; "[red] LECTOTYPUS [printed] Ƥ 2012 DOLERUS PSEUDOANTICUS MALAISE, 1931 [handwritten] M.Heidemaa des." [printed]; " Dolerus (Equidolerus) pseudoanticus Malaise, 1931 Det. M.Heidemaa " [white, framed, printed]. Paralectotypes: 1Ƥ13 from Kamtchatka, NHRS; 1 ovipositor slide, USNM (see taxonomic notes).

Dolerus subfasciatus pseudoanticus: Zhelochovtsev & Zinovjev 1996: 360 .

Distribution. This species is known only by the type series from Kluchi in Kamchatka ( Russia).

Specimens studied. Part of the syntype series (2Ƥ13, see taxonomic notes).

Host plant. Unknown.

Taxonomic notes. A holotype was not designated by Malaise (1931). The syntype series of D. pseudoanticus consists of 4Ƥ and 23 according to the original description. 2Ƥ13 labelled as type (" Typus ", 1Ƥ) and paratypes (1Ƥ [" Paratypus "], 13 ["Allotypus"]) were located at NHRS and studied. A slide preparation of an ovipositor in the USNM, labelled " Dolerus pseudoanticus , Paratype [in fact syntype], female saw, Acc. No. 47009" prepared by Ross was also studied and labelled as paralectotype, but the corresponding specimen was not located. The lancet illustrated by Malaise (1931: fig 2.) fits the lectotype and matches also the paralectotypes. This species was synonymized with D. subfasciatus by Benson (1962), but the two are distinct according to their ovipositor structure (see also Table 1 ). Zhelochovtsev & Zinovjev (1996) treated D. pseudoanticus as a subspecies of D. subfasciatus . A syntype male of D. pseudoanticus was studied and is the only known male of this species (another syntype male was not found). It does not allow assessment of any character variation in the species, but its penis valves differ slightly from D. neoaprilis (see Fig. 17 ).

resembling Equidolerus species with abdominal terga 2–6 unsculptured / weakly sculptured but widely reddish. * The number of

specimens insufficient to assess character variation.

Clypeus emargination mostly 1/2 of the mostly over 1/2 of the 1/2 of the clypeal length 1/2 of the clypeal length

clypeal length clypeal length

Head behind eyes in subparallel / converging parallel / subparallel subparallel subparallel

dorsal view (females)

Figures

FIGURES 4 – 6. Ovipositors / lancet: 4, digital image overlay of D. (E.) neoaprilis lectotype lancet (A) and ovipositor (B) of conspecific non-type specimen (same set of landmarks used as in Fig. 3); 5, D. (E.) pseudoanticus, ovipositor (paralectotype); 6, D. (E.) rhodogaster, ovipositor (lectotype of D. picinus rhodogaster).

FIGURES 11 – 14. Male genitalia: D. (E.) neoaprilis (= D. subfasciatus auct. non F. Smith, USNM): 11, genital capsule in dorsal view; 12, right penis valve in lateral outer aspect; D. (E.) pseudoanticus (paralectotype): 13, genital capsule in dorsal view; 14, right penis valve in lateral outer aspect.

FIGURES 15 – 17. Penis valves (right valves, lateral outer aspect): 15, D. (E.) rhodogaster (non-type, ZIN); 16, D. (E.) lucidus (non-type, BMNH); 17, D. (E.) neoaprilis (non-type, USNM) and D. (E.) pseudoanticus (paralectotype): digital image overlay using landmarks (five encircled crosses correspond to landmarks and black arrows indicate structural details differentiable between D. neoaprilis [smaller arrows] and D. pseudoanticus [larger arrows]).

Tables

TABLE 1. Imaginal diagnostic characters (ordering independent from their diagnostic value) of D. (E.) subfasciatus and three

SPECIES: D. subfasciatus F. Smith CHARACTERS:	D. neoaprilis MacGillivray	D. pseudoanticus Malaise *	D. rhodogaster Zhelochovtsev *
POL / OOCL (in females) 0,54…0,64 (n = 5)	0,50…0,63 (n = 5)	0,48…0,49 (n = 2)	0,56 (n = 1)
Clypeus shape more or less asymmetric	clearly asymmetric	clearly asymmetric	slightly asymmetric